Haplogroup T-M184
Haplogroup T-M184, also known as Haplogroup T is a human Y-chromosome DNA haplogroup. The UEP that defines this clade is the SNP known as M184. Other SNPs – M272, PAGES129, L810, L455, L452, and L445 – are considered to be phylogenetically equivalent to M184. As a primary branch of haplogroup LT, the basal, undivergent haplogroup T* currently has the alternate phylogenetic name of K1b and is a sibling of haplogroup L*. It has two primary branches: T1 and T2.
T-M184 is unusual in that it is both geographically widespread and relatively rare.
T1 – the numerically dominant primary branch of T-M184 – appears to have originated in Western Asia, and possibly spread from there into the East Africa, South Asia, Southern Europe and adjoining regions. T1* may have expanded with the Pre-Pottery Neolithic B culture. Most males who now belong to haplogroup T-L206 carry the subclade T-M70, a primary branch of T-M206. T-M70 nevertheless appears to have long been present in Europe, having possibly arrived there in the Neolithic epoch with the first farmers. This is supported by the discovery of several members of T1a1 at a 7,000 year old settlement, 5800-5400Bc neolithic site in Malak Preslavets, Bulgaria. Autosomal analysis of these remains suggest that some were closely related to modern Southwest Asian populations.
Structure
;Subclade structure of Haplogroup T.- T1
- *T1a
- **T1a1
- ***T1a1a
- ****T1a1a1
- ****T1a1a2
- *****T1a1a2a
- **T1a2
- ***T1a2a
- ***T1a2b
- **T1a3
- ***T1a3a
- ***T1a3b
- T2
Distribution
Overview
Haplogroup T is found at high levels in isolated pockets as far apart as Central Asia, Northeast and Eastern India, Northern Asia, Central Africa, and South Africa. The clade is borne by a majority of Dir clan Somalis in the Horn of Africa; among Kurru, Bauris and Lodha in South Asia; among Toubou in Chad; and in a significant minority of Rajus and Mahli in South Asia; general Somalis, southern Egyptians and Fula in north Cameroon; people from the Chian, Aquilani, Saccensi, Ibizan and Mirandese regions in Europe; Zoroastrians, Bakhtiaris in the Middle East, and Nenets and Kazakhs in Siberia/Central Asia.The maximal worldwide frequency for haplogroup T-M184 is observed among Somalis in the Dire Dawa area and Djibouti, where it accounts for approximately 82% of the Somali male lineages to 100% of the Somali Dir male lineages, respectively. Luis et al. suggest that the presence of T on the African continent may, like R1* representatives, point to an older introduction from Asia. The Levant rather than the Arabian Peninsula appears to have been the main route of entry, as the Egyptian and Turkish haplotypes are considerably older in age than those found in Oman. According to the authors, the spotty modern distribution pattern of haplogroup T-M184 within Africa may therefore represent the traces of a more widespread early local presence of the clade. Later expansions of populations carrying the E1b1b, E1b1a, G and J NRY lineages may have overwhelmed the T-M184 clade-bearers in certain localities.
In the Caucasus and Anatolia it makes up to 4% of the population in southeast and northwest Caucasus as well as in southeast and western Anatolia, peaking up to 20% in Armenians from Sasun. In Middle East it makes up to 4% of the population around the Zagros Mountains and the Persian Gulf as well as around the Taurus Mountains and the Levant basin, peaking up to 10% in Zoroastrians from Kerman, Bakhtiaris, Assyrians from Azerbaijan, Abudhabians, Armenians from Historical Southwestern Armenia and Druzes from Galilee. In Eastern Africa it makes up to 4% of the population on Upper Egypt peaking up to 10% in Luxor.
Haplogroup T is rare almost everywhere in Europe. According to Mendez et al., "the occurrence in Europe of lineages belonging to both T1a1 and T1a2 subclades probably reflects multiple episodes of gene flow. T1a1* haplogroups in Europe likely reflect older gene flow". It makes up to 4% of the population on Central Italy, Western Sicily, Northwest Corsica, Northwestern Iberian Peninsula, Western Andalucia, Western Alps, Eastern Crete, and Macedonia, frequencies up to 10% in Ibiza, Miranda de I Douro, Eastern Oviedo, Cádiz, Badajoz, Balagna, Norma and Ragusa, and peaking at 20% in Sciacca, L'Aquila and some German regions. T-M184 was found in 1.7% of a pool of six samples of males from southwestern Russia, but it was completely absent from a pool of eight samples totalling 637 individuals from the northern half of European Russia. The Russians from the southwest were from the following cities: Roslavl, Livny, Pristen, Repyevka, and Belgorod; and Kuban Cossacks from the Republic of Adygea.
T1 (T-L206)
T1 is the most common descent of T-M184 haplogroup, being the lineage of more than 95% of all Eurasian T-M184 members. One of their descent lineages is found in high frequencies among northern Somali clans. However, it appears to have originated somewhere around the northern Mediterranean Basin, perhaps somewhere between Greece to the Zagros mountains.The basal T1* subclade appears to have spread from northeastern Anatolia, into the Levant at least, with the Pre-Pottery Neolithic B culture. Although it is rare in modern populations, T1* has been found in a Berber individual from Tunisia, a male in Syria, and one sequence among ethnic Macedonians in Macedonia.
T1a (M70)
Mendez et al. points to an ancient presence for T1a-M70 in Europe may reflect early exiles between the ancient lands of Israel and Babylon. The subclade probably arrived with the very first farmers.T1a1*
Recent findings of Haak et al. who discovered several T1a1-CTS880 members in a 7000 years old settlement in Karsdorf, Germany.The T1a1 skeletal remains from this settlement were also found to belong to the H mtdna haplogroup, this settlement has the highest frequency of this mtDNA haplogroup 30.4% that have been found in any early Neolithic Europe population until now.
T1a1, which emerged 17,400-14,600 BP, is the largest lineage downstream from T1a-M70. An individual with T1a1 was first identified in a paper by Tomas et al. in 2006, among a sample of Ibizans from the Balearic Islands of Spain. T-L162 has also been reported in at least one male with a Pontic Greek background.
A subgroup of Ibizans – the Pityusans of the Pityusic Islands – have been found by three different studies to possess T1a1 at relatively high levels of 6.7–16.7%. Tomàs et al. found three cases amongst a sample of 45. Zalloua et al. found nine examples that were L454+ from a sample of 54. Rodriguez et al. found seven cases of L454+ in a sample of 96.
The Pontic Greeks of Anatolia are also reported to possess T1a1. In 2009, a male with the surname Metaxopoulos and a Pontic Greek background was reported to be T-L162 – according to the Y-Chromosome Genome Comparison Project administered by Adriano Squecco. Greeks from the Fatsa reportedly migrated in antiquity from Sinope, which was itself colonised by Ionians. Another ancient Ionian colony in north-west Anatolia, Lámpsakos, had onomastic links to the Pityusic Islands – Lámpsakos was originally an Ionian colony known as Pityussa.
T1a1a (L208)
This lineage, formed 14,200-11,000 BP, is the largest branch downstream T1a1-L162. Firstly discovered and reported at August 2009 in a 23andMe customer of Iberian ancestry that participated in the public Squecco's Y-Chromosome Genome Comparison Project and appearing there as "Avilés" and as "AlpAstur" in 23andMe. Named as "L208" at November 2009.T1a1a1a1b1a1* (T-Y3782*)
One Sardinian male from a sample of 187 – a resident of the Province of Cagliari – has been found to have T-Y3782, also known T1a1a1a1b1a1.T1a1a1a1b1a1a (T-Y3836)
This lineage is mostly found among individuals from the Iberian Peninsula, where the subclade also has its highest diversity. Two subclades can be clearly discriminated. The first, found mainly in post-colonial Puerto Rico, with DYS391=10 and the second, found mainly in Panamá where their Iberian descendants could have the entrance point to America, with DYS439=12.Some members of Y3836 are found among different communities of the Sephardic diaspora but they are found to be extremely rare in the total percentage of some of these communities as seen in Nogueiro et al. This probably could mean that these members could be integrated by these communities through the contact with other native Iberian populations as seen in Monteiro et al. where this lineage was found among native Astur-Leonese speakers.
| Population | Language | Location | Members/Sample size | Percentage | Source | Notes |
| Panamanians | Panamian Castilian | Los Santos Province | 1/30 | 3.3% | ||
| Colombians | Colombian Castilian | Caldas | 2/75 | 2.7% | YHRD | Mestizo individuals |
| Panamanians | Panamian Castilian | Panama Province | 1/43 | 2.3% | ||
| Northwest Argentinians | Argentinian Castilian | Mountainous region of Jujuy | 1/50 | 2% | YHRD | Admixed population |
| Puerto Ricans | Puerto Rican Castilian | Southeast Puerto Rico | 2/110 | 1.8% | ||
| Northeastern Portuguese Jews | Judaeo-Portuguese | Bragança, Argozelo, Carção, Mogadouro, and Vilarinho dos Galegos | 1/57 | 1.8% | ||
| Native Mirandese speakers | Mirandese Astur-Leonese | Miranda de l Douro | 1/58 | 1.7% | ||
| Dominicans | Dominican Castilian | Dominican Republic | 4/261 | 1.5% | ||
| Panamanians | Panamian Castilian | Chiriquí Province | 1/92 | 1.1% | ||
| Mecklenburgers | East Low Saxon | Rostock | 2/200 | 1% | ||
| Mestizos | Colombian Castilian | Bogotá | 2/195 | 1% | YHRD | |
| Mestizos | Colombian Castilian | Valle del Cauca | 1/103 | 1% | YHRD | |
| Mestizos | Ecuadorian Castilian | Quito | 1/102 | 1% | ||
| Venezuelans | Venezuelan Castilian | Maracaibo | 1/111 | 0.9% | ||
| Venezuelans | Venezuelan Castilian | Central Region | 1/115 | 0.9% | ||
| Europeans | Brazilian Portuguese | São Paulo | 1/120 | 0.8 | YHRD | European descents |
| Ecuadorians | Ecuadorian Castilian | Quito | 1/120 | 0.8% | ||
| Colombians | Colombian Castilian | Antioquia | 6/777 | 0.7% | ||
| Mexicans | Mexican Castilian | Mérida | 1/159 | 0.6% | YHRD | Mestizo individuals |
| Eastern Andalusians | Andalusian | Alhama de Granada, Baza, Huéscar, Loja, Montefrío and Órgiva | 1/180 | 0.6% | ||
| Colombians | Colombian Castilian | Santander | 1/193 | 0.5% | YHRD | Mestizo individuals |
| Chileans | Chilean Castilian | Concepción | 1/198 | 0.5% | YHRD | |
| Catalans | Not reported | Metropolitan area of Barcelona | 1/224 | 0.5% | ||
| Mexicans | Mexican Spanish | Guadalajara | 1/246 | 0.4% | YHRD | Mestizo individuals |
| Europeans | Brazilian Portuguese | Rio Grande do Sul | 1/255 | 0.4% |
T2 (PH110)
This lineage could have arrived in the Levant through the PPNB expansion from northeastern Anatolia.A 2014 study found T-PH110 in one ethnic Bhutanese male, out of a sample of 21, possibly implying a rate of 4.8% in Bhutan. Also have been found in a German individual and another two from Caucasus. The Bhutanese and the German haplotypes seems to cluster together.
Possible cases from older research
| Population | Language | Location | Members/Sample size | Percentage | Source | Notes |
| Altaians | Altai | Kurmach-Baygol | 2/11 | 18.2% | K* | |
| Altaians | Altai | Turochak | 2/19 | 10.5% | K | |
| Leoneses | Astur-Leonese | Leon | 1/13 | 7.7% | K | |
| Ossetian Irons | Iron | South Ossetia | 1/21 | 4.8% | No further details available. | |
| Cordobeses | Andalusian | Córdoba | 1/27 | 3.7% | No further details available. | |
| Leoneses | Astur-Leonese | Leon | 2/60 | 3.3% | No further details available. | |
| Tharus | Tharu | Morang | 1/37 | 2.7% | K | |
| Cherkessians | Besleney | Circassia | 2/126 | 1.6% | No further details are available. | |
| Bizkaians | Bizkaiera | Bizkaia | 1/72 | 1.4% | No further details are available. | |
| Europeans | English | Australia | 1/1078 | 0.9% | No further details are available. |
Geographical distribution
Northern Asia
Europe
With K-M9+, unconfirmed but probable T-M70+: 14% of Russians in Yaroslavl, 12.5% of Italians in Matera, 10.3% of Italians in Avezzano, 10% of Tyroleans in Nonstal, 10% of Italians in Pescara, 8.7% of Italians in Benevento, 7.8% of Italians in South Latium, 7.4% of Italians in Paola, 7.3% of Italians in Central-South Italy, 7.1% of Serbs in Serbia, 4.7% of Aromanians in Romania, 3.7% of Italians in Biella, 3.7% of Andalusians in Córdoba, 3.3% of Leoneses in León, 3.2% of Italians in Postua, 3.2% of Italians in Cavaglià, 3.1% of Calabrians in Reggio Calabria, 2.8% of Russians in Ryazan Oblast, 2.8% of Italians in South Apulia, 2.7% of Calabrians in Cosenza, 2.6% of Serbs in Belgrade, 2.5% of Russians in Pskov, 2.4% of Russians in Kaluga, 2.2% of Transylvanians in Miercurea Ciuc, 2.2% of Italians in Trino Vercellese, 1.9% of Italians in Brescia, 1.9% of Romanians in Romania, 1.7% of Serbs and Montenegrins in Serbia and Montenegro, 1.7% of Italians in Marche, 1.7% of Calabrians in Catanzaro, 1.6% of Greeks in Northern Greece, 1.3% of Swiss Germans in Zürich Area, 1.3% of Italians in South Tuscany and North Latium, 1.1% of Dutch in Leiden, 0.5% of Serbs in Novi Sad, 0.5% of Polish in PodlasieOther parts that have been found to contain a significant proportion of haplogroup T-M184 individuals include Trentino, Mariña Lucense, Heraklion, Roslavl, Ourense, Livny, Biella, Entre Douro, Porto, Urbino, Iberian Peninsula, Blekinge/Kristianstad, Belarus, Modena, Provence-Alpes-Côte d'Azur, Pristen, Cáceres, Brac, Satakunta, Western Croatia, Ukrainia, Greifswald, Moldavians in Sofia, Uppsala, Lublin, Pias in Beja, Macedonian Greeks, Nea Nikomedeia, Sesklo/Dimini, Lerna/Franchthi, Açores, Viana do Castelo, Toulouse, Belgorod, Sardinia. According to data from commercial testing, 3.9% of Italian males belonging to this haplogroup. Approximately 3% of Sephardi Jews and 2% of Ashkenazi Jews belong to haplogroup T.
Middle East and Caucasus
Haplogroup T has some significant frequencies in southeast and eastern Anatolia, the Zagros Mountains and both sides of the Persian Gulf.| Population | Language | Location | Members/Sample size | Percentage | Source | Notes |
| Georgians | Georgian | Khashuri | 1/3 | 33.3% | ||
| Priest Zoroastrians | Persian | Shiraz, Tehran and Yazd | 2/8 | 25% | Not specified if Herbad or Mobad | |
| Iraqi Jews | Judeo-Iraqi Arabic | Iraq | 7/32 | 21.9% | 12.5% T1a1a1a1a1a1-P77 and 9.4% T1a3-Y11151 | |
| Armenian Sasuntzis | Western Armenian dialect, Kurmanji and Dimli languages | Sasun | 21/104 | 20.2% | T1a1 and T1a2 subclades | |
| Georgians | Georgian | Sighnaghi and Gurjaani | 2/10 | 20% | ||
| Georgians | Georgian | Kharagauli | 1/5 | 20% | ||
| Kumyks | Kumyk | Daghestani lowlands | 2/10 | 20% | Reported as K* but according to Karafet16 and Yunusbayev12 only T fits. | |
| Kurdish Jews | Judeo-Aramaic | Kurdistan | 19/99 | 19.2% | ||
| Kurdish Jews | Judeo-Aramaic | Kurdistan | 9/50 | 18% | 10% T1a1a1a1a1a1-P77 and 8% T1a1-L162 | |
| Druzes | Palestinian Arabic | Galilee | 7/40 | 17.5% | ||
| Assyrians | Aramaic | refugees in Armenia | 16/106 | 15.1% | Reported as K*. Their homeland in the areas around Urmia. | |
| Assyrians | Aramaic | Unknown | 4/28 | 14.3% | ||
| Georgians | Georgian | Dusheti | 1/7 | 14.3% | ||
| Iranian Jews | Judeo-Iranian | Iran | 3/22 | 13.6% | 4.5% T1a1a1a1a1a1-P77 and 9.1% T1a3-Y11151 | |
| Zoroastrians | Persian | Kerman | 5/37 | 13.5% | ||
| Iraqi Jews | Judeo-Iraqi Arabic | Iraq | 13/99 | 13.1% | ||
| Bakhtiaris | Bakhtiari | Izeh | 13/103 | 12.6% | ||
| Mountain Jews | Judeo-Tat | Derbentsky District | 2/17 | 11.8% | All belong to T1a1a1a1a1a1-P77 | |
| Armenians | Western Armenian dialect | Historical Southwestern Armenia | 11/96 | 11.5% | ||
| Abudhabians | Gulf Arabic | Abu Dhabi | 21/191 | 11% | ||
| Assyrians | Assyrian | West Azerbaijan Province | 4/39 | 10.3% | ||
| Iranian Jews | Judeo-Iranian | Iran | 5/49 | 10.2% | ||
| Persian Muslims | Persian | Shiraz | 5/51 | 9.8% | ||
| Persian Muslims | Persian | Kerman | 6/66 | 9.1% | ||
| Iraqis | Iraqi Arabic | Al-Qadisiyah | 6/69 | 8.7% | ||
| Armenians | Armenian | Armenia | 35/413 | 8.5% | ||
| Kurds | Sorani | Kurdestan | 5/59 | 8.5% | ||
| Omani Arabs | Omani Arabic | Oman | 10/121 | 8.3% | ||
| Kurds | Sorani | Kurdestan | 2/25 | 8% | ||
| Azeris | Azeri | West Azerbaijan Province | 5/63 | 7.9% | ||
| Mazanderanis | Mazanderan | Mazandaran | 1/13 | 7.7% | ||
| Cypriots | Cypriot Greek | Cyprus | 3/41 | 7.3% | ||
| Iraqis | Iraqi Arabic | Iraq | 10/139 | 7.2% | ||
| Kuwaitis | Gulf Arabic | Kuwait | 3/42 | 7.1% | ||
| Iraqis | Iraqi Arabic | Iraq | 3/43 | 7% | ||
| Arabs | Levantine Arabic | Israel and Palestine | 10/143 | 7% | ||
| Persians | Farsi | Fars | 3/44 | 6.8% | ||
| Christian Arabs | Levantine Arabic | Israel and Palestine | 3/44 | 6.8% | ||
| Western Armenians | Armenian | Eastern Turkey | 6/90 | 6.7% | ||
| Persians | Farsi | Yazd | 3/46 | 6.5% | ||
| Armenians | Armenian | Gardman | 6/96 | 6.3% | ||
| Yezidis | Kurmanji | refugees in Armenia | 12/196 | 6.1% | Reported as K*. Their homeland in the areas around Laliş. | |
| Muslim Arabs | Levantine Arabic | Israel and Palestine | 7/119 | 5.9% | ||
| Zahedan, Baluchestan, Iran | 6/103 | 5.8% | ||||
| Northern Armenians | Armenian | Northern Armenia, southern Georgia and northwestern Azerbaijan | 10/189 | 5.3% | ||
| Armenians | Armenian | Tehran | 2/38 | 5.3% | ||
| Eastern Armenians | Armenian | Karabakh | 11/215 | 5.1% | ||
| Persians | Farsi | Khorasan | 3/59 | 5.1% | ||
| Saudi Arabians | Arabic dialects | Saudi Arabia | 8/157 | 5.1% | ||
| Armenians | Armenian | Syunik | 7/140 | 5% | ||
| Emiratis | Gulf Arabic | United Arab Emirates | 8/164 | 4.9% | ||
| Lebanese Muslims | Lebanese Arabic | Lebanon | 28/568 | 4.9% | ||
| Cypriots | Cypriot Greek | Lemesos | 6/126 | 4.8% | ||
| Kumyks | Kumyk | Khasavyurtovsky District | 1/21 | 4.8% | ||
| Avars | Avar | southeastern Dagestan | 2/42 | 4.8% | ||
| Kurds | Kurmanji | Anatolia | 12/251 | 4.8% | ||
| Kurds | Kurdish dialects | Kurdistan | 6/126 | 4.8% | ||
| Anizes | Gulf Arabic | Kuwait | 1/21 | 4.7% | ||
| Lebaneses | Levantine Arabic | Lebanon | 43/914 | 4.7% | ||
| Cypriots | Cypriot Greek | Cyprus | 3/65 | 4.6% | ||
| Maronites | Lebanese Arabic and Syriac | Lebanon | 24/518 | 4.6% | ||
| Armenians | Armenian | Ararat | 2/44 | 4.6% | ||
| Muslim Kurds | Kurdish dialects | Kurdistan | 4/95 | 4.2% | ||
| Qeshmis | Qishmi | Qeshm | 2/49 | 4.1% | ||
| Lurs | Luri | Lorestan | 2/50 | 4% | ||
| Sadats | Languages of Iran | Different cities of Iran | 2/50 | 4% | ||
| Persians | Persian | Eastern Iran | 3/77 | 3.9% | ||
| Armenians | Armenian | Lake Van | 4/103 | 3.9% | ||
| Saudi Arabians | Arabic dialects | Saudi Arabia | 4/106 | 3.8% | ||
| Turkish Cypriots | Cypriot Turkish | 138 different villages, towns or cities from Cyprus | 14/380 | 3.7% | Paternal lineages originating from the traditional Turkish Cypriot settlements throughout the island | |
| Birjand, South Khorasan, Iran | 1/27 | 3.7% | All T1a3-Y12871 | |||
| Armenians | Armenian | Ararat Valley | 4/110 | 3.6% | ||
| Armenians | Armenian | Armenia | 2/57 | 3.5% | ||
| Georgians | Georgian | Omalo | 1/29 | 3.5% | ||
| Iranians | Languages of Iran | South Iran | 4/117 | 3.4% | ||
| Ionians | Greek | Phokaia | 1/31 | 3.2% | ||
| Bandaris | Bandari | Bandar Abbas | 4/131 | 3.1% | ||
| Cypriots | Cypriot Greek | Larnaka | 2/67 | 3% | ||
| Alans | Karachay-Baksan-Chegem | Kabardino-Balkaria | 1/69 | 2.9% | ||
| Jordanians | Arabic dialects | Jordania | 8/273 | 2.9% | ||
| Cypriots | Cypriot Greek | Ammochostos | 3/122 | 2.5% | ||
| Lezghins | Lezgian | Southern Dagestan | 2/81 | 2.5% | ||
| Turks | Turkish | Turkey | 13/523 | 2.5% | ||
| Persians | Persian | Esfahan | 1/13 | 2.4% | ||
| Iranians | Languages of Iran | Iran | 7/324 | 2.2% | ||
| Azerbaijani Muslims | Azerbaijani | Uromia | 2/91 | 2.2% | ||
| Yemenite Jews | Hebrew and Arabic | Yemen | 2/94 | 2.1% | ||
| Andis | Andi | western Dagestan | 1/49 | 2% | ||
| Cypriots | Cypriot Greek | Paphos | 2/105 | 1.9% | ||
| Cypriots | Cypriot Greek | Nicosia | 3/161 | 1.9% | ||
| Assyrians | Assyrian Neo-Aramaic | Uromia and Tehran | 1/55 | 1.8% | ||
| Abkhazians | Abkhaz | Abkhazia | 1/58 | 1.7% | ||
| Kuwaitis | Gulf Arabic | Kuwait | 2/117 | 1.7% | ||
| Greek Orthodox | Koine Greek | Lebanon | 2/116 | 1.7% | ||
| Mashhad, Razavi Khorasan, Iran | 2/129 | 1.6% | 0.8% T1a3-Y11151 | |||
| Aeolians | Greek | Smyrna | 1/68 | 1.5% | ||
| Georgians | Georgian | Georgia | 1/66 | 1.5% | ||
| Turkmens | Turkmen | Golestan | 1/68 | 1.5% | ||
| Kumyks | Kumyk | Northern Dagestan | 1/73 | 1.4% | ||
| Kuban Nogays | Nogai | north of Sea of Azov around Prymorsk | 1/87 | 1.2% | ||
| Ossetian Digors | Digorian | North Ossetia | 1/127 | 0.8% | ||
| Yemeni Arabs | Sanaani Arabic | Sana'a | 1/129 | 0.8% | ||
| Syrians | Syrian Arabic | Syria | 4/518 | 0.8% | ||
| Kabardins | Kabardian | Kabardino-Balkaria | 1/140 | 0.7% | ||
| Circassians | Adyghe | Republic of Adygea | 1/142 | 0.7% | ||
| Abkhazians | Abkhaz | Abkhazia | 1/162 | 0.6% |
There are also unconfirmed reports of T-M70+ amongst 28% of Lezginians in Dagestan, 21.7% of Ossetians in Zamankul, 14% of Iranians in Isfahan, 13% of Ossetians in Zil'ga, 12.6% of Kurmanji Kurds in Eastern Turkey, 11.8% of Palestinian Arabs in Palestine, 8.3% of Iranians in Shiraz, 8.3% of Ossetians in Alagir, 8% of Kurmanji Kurds in Georgia, 7.5% of Iranians in Tehran, 7.4% of Palestinian Arabs in Israeli Village, 7% of Palestinian Arabs in Israel and Palestine, 5% of Chechens in Chechenia, 4.2% of Azerbaijanians in Azerbaijan, 4.1% of Iranians in Isfahan, 4% of Armenians in Armenia, 4% of Bedouins in Israel and 2.6% of Turks in Ankara.
Africa
Fossils excavated at the Late Neolithic site of Kelif el Boroud in Morocco, which have been radiocarbon-dated to around 3,000 BCE, have been found to belong to haplogroup T-M184.| Population | Language | Location | Members/Sample size | Percentage | Source | Notes |
| Somalis | Somali | Djibouti | 24/24 | 100% | The main sub-clans of the Dir clan in Djibouti are the Issa and Gadabuursi. | |
| Somalis | Somali | Dire Dawa | 14/17 | 82.4% | Dir sub-clans of Dire Dawa are Issa, Gurgura and Gadabuursi. | |
| Anteony | Antemoro | old Antemoro Kingdom | 22/37 | 59.5% | The Anteony are the descendants of aristocrats, from whom the Antemoro king is chosen. Can be grouped into the Silamo, because they have the right to undertake the ritual slaughter of animals | |
| Somalis and Afars | Somali and Afar | Djibouti | 30/54 | 56.6% | Mixed sample of Somali and Afar individuals. | |
| Somalis | Somali | Shilavo | 5/10 | 50% | The geographic location of this Ethiopia sample as seen in Fig.1. | |
| Toubou | Toubou | Chad | 31% | All belonging to the T1a-PF5662 subclade | ||
| Afars | Afar language | Djibouti | 5/20 | 25% | ||
| Akie | Akie people | Tanzania | 3/13 | 23.1% | Akie people have remnants of a Cushitic language | |
| Somalis | Somali | Jijiga | 19/83 | 22.9% | Jijiga Somalis. | |
| Arabs from Somalia | Somali | immigrants in Yemen | 7/33 | 21.2% | ||
| Lemba | Venda and Shona | South Africa | 6/34 | 17.6% | Exclusively belong to T1a2*. Possible recent founder effect. Low frequency of T1a2 has been observed in Bulgarian Jews and Turks but is not found in other Jewish communities. Y-str Haplotypes close to some T1a2 Armenians. | |
| Rangi | Rangi Language | Tanzania | 5/32 | 15.6% | ||
| - | Somalia | 15/105 | 14.3% | - | ||
| Iraqw | Iraqw language | Tanzania | 6/47 | 12.8% | ||
| Wachagga | Kichagga | Dār as-Salām | 3/24 | 12.5% | Mixed with Rift Southern Cushites. | |
| Somali | Somali | immigrants to Norway | 12/104 | 11.5% | ||
| Bench | Bench | Bench Maji Zone | 14/126 | 11.4% | ||
| Kores | SNNP | 2/18 | 11.1% | |||
| Oromo | Afaan Oromo language | Oromiyaa | 1/9 | 11.1% | ||
| Fulbe | Fula | northern Cameroon | 3/27 | 11.1% | ||
| Gorowa | Gorowa language | Tanzania | 2/19 | 10.5% | ||
| Somali | Somali | immigrants to Denmark | 21/201 | 10.4% | ||
| Upper Egyptians | Egyptian Arabic | Luxor Governorate | 3/29 | 10.3% | ||
| Kontas | Konta language | Konta special woreda | 11/107 | 10.3% | ||
| Rendille | Rendille language | Marsabit County | 3/31 | 9.7% | ||
| Datogs | Rendille language | Tanzania | 3/31 | 9.7% | ||
| Gewadas | Gewada language | SNNP | 11/116 | 9.5% | ||
| Antalaotra | Antemoro | old Antemoro Kingdom | 4/43 | 9.3% | The Antalaotra are in charge of the magical and religious domains; they have the ability to read and write Sorabe. Can be grouped into the Silamo, because they have the right to undertake the ritual slaughter of animals | |
| Upper Egyptians | Egyptian Arabic | Aswan Governorate | 1/11 | 9.1% | ||
| N’Djamena Mix | Mix | N’Djamena | 5/55 | 9.1% | Marc Haber 2016 | All belonging to the T1a-PF5662 subclade |
| Upper Egyptians | Egyptian Arabic | Assiut Governorate | 6/70 | 8.6% | ||
| Konsos | Konso special woreda | 2/24 | 8.3% | |||
| Somali | Somali | immigrants to Sweden | 12/147 | 8.2% | ||
| Arabs and Berbers | Egyptian Arabic and Siwi | Lower Egypt | 12/147 | 8.2% | ||
| Upper Egyptians | Egyptian Arabic | Sohag Governorate | 4/52 | 7.7% | ||
| Egyptians | Erythraic | Egypt | 7/92 | 7.6% | If the K* sample is M184+ then 8.7% | |
| Tigrayans | Tigrinya | Tigray Region | 2/30 | 6.7% | ||
| Dirashas | Dirasha | Dirashe special woreda | 5/79 | 6.3% | ||
| Canarians | Canarian Spanish | Tenerife | 11/178 | 6.2% | ||
| Kordofanians | Kordofanian | Kurdufan | 4/69 | 5.8% | ||
| Upper Egyptians | Egyptian Arabic | Qena Governorate | 3/52 | 5.8% | ||
| Tuareg | Tuareg | Gorom-Gorom | 1/18 | 5.6% | ||
| Afars | Afar | Afar Region | 6/111 | 5.4% | ||
| Ethiopians | Ethiopian languages | Ethiopia | 4/74 | 5.4% | ||
| Mashiles | Mashile language | SNNP | 7/130 | 5.4% | ||
| Gurages | Gurage languages | SNNP | 6/118 | 5.1% | ||
| Turu | Nyaturu | Tanzania | 1/20 | 5% | ||
| Moroccan Jews | Haketia | Israel | 1/20 | 5% | ||
| Gedeos | Gedeo | SNNP | 6/122 | 4.9% | ||
| Wairak | Iraqw | Tanzania | 2/41 | 4.9% | ||
| Western Libyans | Libyan Arabic | Tripoli region | 7/142 | 4.9% | ||
| Tunisians | Tunisian Arabic | Sfax | 5/105 | 4.8% | ||
| Libyans | Libyan Arabic | Tripoli area | 3/63 | 4.8% | ||
| Kanuri | Kanuri | Cameroon | 1/21 | 4.8% | ||
| Iraqw | Iraqw | Tanzania | 2/43 | 4.7% | ||
| Yems | Yemsa | SNNP | 5/107 | 4.7% | ||
| Jews | Ethiopia | 1/22 | 4.5% | |||
| Gobeze | Cushitic | SNNP | 5/113 | 4.4% | ||
| Upper Egyptians | Egyptian Arabic | Minya Governorate | 1/23 | 4.3% | ||
| Konsos | Konso language | Konso special woreda | 4/94 | 4.3% | ||
| Kembaatas | East Cushitic | Kembata Tembaro Zone | 4/102 | 3.9% | ||
| Tigrayans | Tigrinya | Eritrea | 1/28 | 3.6% | ||
| Tigrayans | Tigrinya | Eritrea | 1/31 | 3% | ||
| Amharas | Amharic | Ethiopia | 1/34 | 2.9% | ||
| Hutus | Rwanda-Rundi | Rwanda | 1/39 | 2.6% | ||
| Lower Egyptians | Egyptian Arabic | Mansoura | 1/44 | 2.2% | ||
| Berbers | Shilha | Siwa Oasis | 2/93 | 2.2% | ||
| Berbers | Jerba Berber | Djerba | 1/47 | 2.1% | ||
| Meru | Meru | Tanzania | 2/99 | 2% | ||
| Itam | Ibibio | Obong Itam | 1/50 | 2% | ||
| Cape Verdeans | Cape Verdean Creole | Windward islands São Nicolau, São Vicente, and Santo Antão | 2/101 | 2% | ||
| Ovimbundo | Umbundu and Portuguese | Angola | 1/53 | 1.9% | ||
| Tunisians | Tunisian Arabic | Tunis | 1/54 | 1.9% | ||
| Berbers | Shilha | Asni | 1/54 | 1.9% | ||
| Eastern Libyans | Libyan Arabic | Benghazi | 4/214 | 1.9% | ||
| Algerians | Algerian Arabic | Algeria | 3/164 | 1.8% | ||
| Baribas | Baatonum | Benin | 1/57 | 1.8% | T1a-M70 | |
| Bokoras | Karamojong | Karamoja region | 1/59 | 1.7% | ||
| Lower Egyptians | Egyptian Arabic | Cairo | 1/63 | 1.6% | ||
| Tumbuka | Tumbuka | northern Malawi | 1/61 | 1.6% | ||
| Mozabites | Mozabite | Ghardaia | 1/68 | 1.5% | ||
| Tunisians | Tunisian Arabic | South Tunisia | 3/200 | 1.5% | ||
| Soussians | Tunisian Arabic | Sousse | 3/220 | 1.4% | ||
| Chewa | Chewa | Malawi | 1/92 | 1.1% | ||
| Maasai | Maasai | Kinyawa | 1/100 | 1% | YHRD | |
| Bantu | Narrow Bantu | Pretoria | 1/98 | 1% | ||
| Nilotes | Ateker | Karamoja region | 1/118 | 0.8% | ||
| Andalusians | Andalusian Arabic | Testour, El Alia, Gualaat-El-Andalous, Slouguia | 1/132 | 0.8% | Refugees from Al-Andalus following the capitulation of the Islamic kingdoms in Valencia and Granada | |
| Bantus | Bantu | Botswana, Namibia and Zambia | 1/140 | 0.7% | Father and paternal grandfather belonged to the same ethnolinguistic group | |
| Basothos | Sesotho | Lesotho | 1/181 | 0.6% | ||
| Moroccans | Moroccan Arabic | Casablanca metropolitan area | 1/166 | 0.6% | The industrial capital of Morocco where the urban growth is maintained by immigration from all parts of Morocco | |
| Khoisans | Khoisan | Botswana, Namibia and Zambia | 1/371 | 0.3% | Father and paternal grandfather belonged to the same ethnolinguistic group |
South Asia
Haplogroup T-M184 has been detected at very high levels in some parts of eastern India.T1a-M70 in India has been considered to be of West Eurasian origin.
| Population | Language | Location | Members/Sample size | Percentage | Source | Notes |
| Kurru | Yerukala | Andhra Pradesh | 10/18 | 55.6% | ||
| Bauris | Bengali | West Bengal | 10/19 | 52.6% | K* is found at 6/19, if M70- but M184+, then could be 84.2%. Bauris are thought to be descendants of a native tribe of the Central Highlands before the Aryan invasion, then as Bauris have not been well assimilated and have not participated satisfactorily in the new Aryan society, the Bauris ended up being seen as "low caste". They are at "halfway" between the old Bauri tribal and the new Aryan society lifestyle. | |
| Lodha | Lodhi | West Bengal | 2/4 | 50% | ||
| Rajus | Telugu | Andhra Pradesh | 3/19 | 15.9% | ||
| Maheli | Mahali | West Bengal | 2/13 | 15.3% | ||
| Chenchus | Chenchu | Andhra Pradesh | 3/20 | 15% | K* is found at 7/20, if M70- but M184+, then could be 50% | |
| Kare Vokkal | Kannada | Uttara Kannada | 4/30 | 13.3% | K* is found at 3/30, if M70- but M184+, then could be 23.3% | |
| Banjaras | Lambadi | Andhra Pradesh | 2/18 | 11.1% | ||
| Gonds | Gondi | South Uttar Pradesh | 4/38 | 10.6% | ||
| Gonds | Gondi | Madhya Pradesh | 10/139 | 7.2% | ||
| Indians | languages of India | South India | 18/305 | 5.9% | ||
| Maheli | Mahali | Jamshedpur from Jharkhand; Purulia, Midnapore & other location from West Bengal | 2/38 | 5.3% | Two samples from different studies grouped together | |
| Chenchus | Chenchu | Andhra Pradesh | 3/61 | 4.9% | Samples from Trivedi et al. and Kivisild et al. | |
| Banjaras | Lambadi | Andhra Pradesh | 2/53 | 3.8% | Two samples from different studies grouped together | |
| Indians | languages of India | East India | 14/367 | 3.8% | ||
| Gujaratis | Gujarati | Gujarat | 1/29 | 3.4% | ||
| Lodha | Lodhi | Midnapore & other location from West Bengal | 2/71 | 2.8% | Three samples from different studies grouped together | |
| Sahariyas | Saharia | Madhya Pradesh | 2/73 | 2.7% | ||
| Tamtas | Bageshwar | 1/34 | 2.9% | |||
| Kshatriyas | Pithoragarh | 2/79 | 2.5% | |||
| Aryas | Arya | Nainital | 1/46 | 2.2% | ||
| Laotians | Lao | Laos | 1/53 | 1.9% | ||
| Maravars | Tamil | Ramanathapuram | 1/80 | 1.3% | Dry Land Farmers | |
| Garos | Garo | Tangail | 1/120 | 0.8% | Likely P77+ |
With K-M9+, unconfirmed but probable T-M70+: 56.6% of Kunabhis in Uttar Kannada, 32.5% of Kammas in Andhra Pradesh, 26.8% of Brahmins in Visakhapatnam, 25% of Kattunaiken in South India, 22.4% of Telugus in Andhra Pradesh, 20% of Ansari in South Asia, of Poroja in Andhra Pradesh, 9.8% of Kashmiri Pandits in Kashmir, 8.2% of Gujars in Kashmir, 7.7% of Siddis in Andhra Pradesh, 5.5% of Adi in Northeast India, 5.5% of Pardhans in Adilabad, 5.3% of Brahmins in Bihar, 4.3% of Bagata in Andhra Pradesh, 4.2% of Valmiki in Andhra Pradesh, of Brahmins in Maharashtra, 3.1% of Brahmins in Gujarat, 2.9% of Rajput in Uttar Pradesh, 2.3% of Brahmins in Peruru, and 1.7% of Manghi in Maharashtra.
Also in Desasth-Brahmins in Maharashtra and Chitpavan-Brahmins in Konkan, Chitpavan-Brahmins in Konkan.
Central Asia & East Asia
Unconfirmed but probable T-M70+: 2% of Hui in Liaoning, and 0.9% of Bidayuh in Sarawak.Americas (post-colonisation)
Ancient DNA
Ancient DNA from 'Ain Ghazal
| Ain Ghazal PPNB individual | Ghazal-I I1707 AG83_5 Poz-81097 |
| Y DNA | T1-PF5610 |
| Population | Neolithic Farmers |
| Language | |
| Inferred cultural affiliation | Late Middle PPNB |
| Date | 9573 ± 39 |
| House/location | Ain Ghazal |
| Number | 1/2 |
| Percentage | 50% |
| mtDNA | R0a |
| Isotope Sr | |
| Eye color | Likely non-Dark |
| Hair color | Likely non-Dark |
| Skin pigmentation | Light |
| ABO Blood Group | Likely O or B |
| Diet | |
| FADS activity | rs174551, rs174553, rs174576 |
| Lactose intolerance | Likely lactose-intolerant |
| DNA shared with Oase-1 | 14.2% |
| DNA shared with Ostuni1 remains | 6.7% |
| SDNA shared with Neanderthal Vi33.26 | 0.93% |
| DNA shared with Neanderthal Vi33.25 | 1.2% |
| DNA shared with Neanderthal Vi33.16 | 0.3% |
| Ancestral components | Neolithic Anatolia/Southeast Europe: 56.82%, Paleolithic Levant : 24.09%, Caucasus Hunter / Early European Farmer: 12.51%, Scandinavian / West European Hunter: 4.16%, Sub Saharan: 2.04%, East European Hunter: 0.37% |
| puntDNAL K12 Ancient | |
| Dodecad | |
| Eurogenes | |
| Dodecad | |
| Genetic distance | |
| Parental consanguinity | |
| Age at death | |
| Death position | |
| SNPs | 152.234 |
| Read Pairs | |
| Sample | |
| Source | Lazaridis 2016 |
| Notes | Evidence of a northerly origin for this population, possibly indicating an influx from the region of northeastern Anatolia. |
Haplogroup T is found among the later middle Pre-Pottery Neolithic B inhabitants from the 'Ain Ghazal archaeological site. It was not found among the early and middle PPNB populations. It is thought that the Pre-Pottery Neolithic B population is mostly composed of two different populations: members of early Natufian civilisation and a population resulting from immigration from the north, i.e. north-eastern Anatolia. However, Natufians have been found to belong mostly to the E1b1b1b2 lineage – which is found among 60% of the whole PPNB population and 75% of the 'Ain Ghazal population, being present in all three middle PPNB stages.
Later middle PPNB populations in the Southern Levant were already witnessing severe changes in climate that would have been exacerbated by large population demands on local resources. Beginning at 8.9 cal ka BP we see a significant decrease in population in highland Jordan, ultimately leading to the complete abandonment of almost all central settlements in this region.
The 9th millennium Pre-Pottery Neolithic B period in the Levant represents a major transformation in prehistoric lifeways from small bands of mobile hunter–gatherers to large settled farming and herding villages in the Mediterranean zone, the process having been initiated some 2–3 millennia earlier.
'Ain Ghazal is situated in a relatively rich environmental setting immediately adjacent to the Wadi Zarqa, the longest drainage system in highland Jordan. It is located at an elevation of about 720m within the ecotone between the oak-park woodland to the west and the open steppe-desert to the east.
Evidence recovered from the excavations suggests that much of the surrounding countryside was forested and offered the inhabitants a wide variety of economic resources. Arable land is plentiful within the site's immediate environs. These variables are atypical of many major neolithic sites in the Near East, several of which are located in marginal environments. Yet despite its apparent richness, the area of 'Ain Ghazal is climatically and environmentally sensitive because of its proximity throughout the Holocene to the fluctuating steppe-forest border.
The Ain Ghazal settlement first appear in the middle PPNB, which is split into two phases. Phase 1 starts 10300 yBP and ends 9950 yBP, phase 2 ends 9550 yBP.
The estimated population of the middle PPNB site from ‘Ain Ghazal is of 259-1,349 individuals with an area of 3.01-4.7 ha. Is argued that at its founding at the commencement of the middle PPNB ‘Ain Ghazal was likely 2 ha in size and grew to 5 ha by the end of the middle PPNB. At this point in time their estimated population was 600-750 people or 125-150 people per hectare.
Notable haplogroup members
Elite endurance runners
Possible patterns between Y-chromosome and elite endurance runners were studied in an attempt to find a genetic explanation to the Ethiopian endurance running success. Given the superiority of East African athletes in international distance running over the past four decades, it has been speculated that they are genetically advantaged. Elite marathon runners from Ethiopia were analysed for K* which according to the previously published Ethiopian studies is attributable to the haplogroup TAccording to further studies, T1a1a* was found to be proportionately more frequent in the elite marathon runners sample than in the control samples than any other haplogroup, therefore this y-chromosome could play a significant role in determining Ethiopian endurance running success. Haplogroup T1a1a* was found in 14% of the elite marathon runners sample of whom 43% of this sample are from Arsi province. In addition, haplogroup T1a1a* was found in only 4% of the Ethiopian control sample and only 1% of the Arsi province control sample. T1a1a* is positively associated with aspects of endurance running, whereas E1b1b1 is negatively associated.
Thomas Jefferson
A notable member of the T-M184 haplogroup is American President Thomas Jefferson. The Y-chromosomal complement of the Jefferson male line was studied in 1998 in an attempt to resolve the controversy over whether he had fathered the mixed-race children of his slave Sally Hemings. A 1998 DNA study of the Y chromosome in the Jefferson male line found that it matched that of a descendant of Eston Hemings, Sally Hemings' youngest son. This confirmed the body of historical evidence, and most historians believe that Jefferson had a long-term intimate liaison with Hemings for 38 years, and fathered her six children of record, four of whom lived to adulthood. In addition, the testing conclusively disproved any connection between the Hemings descendant and the Carr male line. Jefferson grandchildren had asserted in the 19th century that a Carr nephew had been the father of Hemings' children, and this had been the basis of historians' denial for 180 years.Jefferson's paternal family traced back Wales, where T is incredibly rare, as it is throughout Britain. A couple of British males with the Jefferson surname have been found with the third president's type of T, reinforcing the idea that his immediate paternal ancestry was British.
Phylogenetic tree
Nomenclatural history
Prior to 2002, there were in academic literature at least seven naming systems for the Y-Chromosome Phylogenetic tree. This led to considerable confusion. In 2002, the major research groups came together and formed the Y-Chromosome Consortium. They published a joint paper that created a single new tree that all agreed to use. Later, a group of citizen scientists with an interest in population genetics and genetic genealogy formed a working group to create an amateur tree aiming at being above all timely. The table below brings together all of these works at the point of the landmark 2002 YCC Tree. This allows a researcher reviewing older published literature to quickly move between nomenclatures.| YCC 2002/2008 | YCC 2002 | YCC 2005 | YCC 2008 | YCC 2010r | ISOGG 2006 | ISOGG 2007 | ISOGG 2008 | ISOGG 2009 | ISOGG 2010 | ISOGG 2011 | ISOGG 2012 | ISOGG 2013 | |||||||
| T-M184 | 26 | VIII | 1U | 25 | Eu16 | H5 | F | K* | K | T | T | K2 | K2 | T | T | T | T | T | T |
| K-M70/T-M70 | 26 | VIII | 1U | 25 | Eu15 | H5 | F | K2 | K2 | T | T1 | K2 | K2 | T | T | T | T1 | T1a | T1a |
| T-P77 | 26 | VIII | 1U | 25 | Eu15 | H5 | F | K2 | K2 | T2 | T1a2 | K2 | K2 | T2 | T2 | T2a1 | T1a1b | T1a1a1 | T1a1a1 |
Original research publications
The following research teams per their publications were represented in the creation of the YCC Tree.α and
β
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η