Phlegmariurus
Phlegmariurus is a genus of lycophyte plants in the family Lycopodiaceae. The genus is recognized in the Pteridophyte Phylogeny Group classification of 2016, but not by some other sources, which keep it in a broadly defined Huperzia.
Taxonomy
The genus was first described in 1909 by Wilhelm Herter as the section Phlegmariurus of the genus Lycopodium. The section was elevated to a genus by Josef Ludwig Holub in 1964.Within the family Lycopodiaceae, Phlegmariurus is placed in the subfamily Huperzioideae. A phylogenetic study in 2016, employing both molecular and morphological data, concluded that either a one-genus or a three-genus division of the subfamily produced monophyletic taxa. The authors preferred the three-genus division, recognizing Huperzia, Phlegmariurus and Phylloglossum. Their preferred hypothesis for the relationships of the three genera was:
The majority of the species formerly placed in a broadly defined Huperzia belong in Phlegmariurus. However, the genera are difficult to separate morphologically, and others have preferred the one-genus division of the subfamily.
Species
, the Checklist of Ferns and Lycophytes of the World accepted over 300 species:- Phlegmariurus acerosus B.Øllg.
- Phlegmariurus acifolius B.Øllg.
- Phlegmariurus acutus B.Øllg.
- Phlegmariurus affinis B.Øllg.
- Phlegmariurus afromontanus Pic. Serm.
- Phlegmariurus ambrensis A.R.Field & Bauret
- Phlegmariurus amentaceus B.Øllg.
- Phlegmariurus andinus B.Øllg.
- Phlegmariurus apolinari-mariae comb. ined.
- Phlegmariurus aqualupianus B.Øllg.
- Phlegmariurus arcturi B.Øllg.
- Phlegmariurus arcuatus B.Øllg.
- Phlegmariurus aristei comb. ined.
- Phlegmariurus ascendens B.Øllg.
- Phlegmariurus attenuatus B.Øllg.
- Phlegmariurus australis A.R.Field
- Phlegmariurus austroecuadoricus B.Øllg.
- Phlegmariurus austrosinicus Li Bing Zhang
- Phlegmariurus axillaris comb. ined.
- Phlegmariurus badinianus B.Øllg.
- Phlegmariurus balansae A.R.Field & Bostock
- Phlegmariurus bampsianus A.R.Field & Bostock
- Phlegmariurus banayanicus A.R.Field & Bostock
- Phlegmariurus beitelianus comb. ined.
- Phlegmariurus beitelii B.Øllg.
- Phlegmariurus biformis B.Øllg.
- Phlegmariurus binervius B.Øllg.
- Phlegmariurus bolanicus A.R.Field & Bostock
- Phlegmariurus brachiatus B.Øllg.
- Phlegmariurus brachystachys A.R.Field & Bostock
- Phlegmariurus bradeorum B.Øllg.
- Phlegmariurus brassii A.R.Field & Bostock
- Phlegmariurus brevifolius B.Øllg.
- Phlegmariurus brongniartii B.Øllg.
- Phlegmariurus buesii B.Øllg.
- Phlegmariurus callitrichifolius B.Øllg.
- Phlegmariurus campianus B.Øllg.
- Phlegmariurus cancellatus Ching
- Phlegmariurus capellae B.Øllg.
- Phlegmariurus capillaris B.Øllg.
- Phlegmariurus carinatus Ching
- Phlegmariurus catacachiensis B.Øllg.
- Phlegmariurus catharinae comb. ined.
- Phlegmariurus cavifolius A.R.Field & Bostock
- Phlegmariurus ceylanicus comb. ined.
- Phlegmariurus chamaeleon B.Øllg.
- Phlegmariurus changii T.Y.Hsieh
- Phlegmariurus chiricanus B.Øllg.
- Phlegmariurus christii B.Øllg.
- Phlegmariurus ciliatospiculatus B.Øllg.
- Phlegmariurus ciliolatus B.Øllg.
- Phlegmariurus clarkianus comb. ined.
- Phlegmariurus cleefianus B.Øllg.
- Phlegmariurus colanensis B.Øllg.
- Phlegmariurus columnaris B.Øllg.
- Phlegmariurus comans comb. ined.
- Phlegmariurus compactus B.Øllg.
- Phlegmariurus copelandianus A.R.Field
- Phlegmariurus coralium A.R.Field & Bostock
- Phlegmariurus coriacea comb. ined.
- Phlegmariurus costaricensis comb. ined.
- Phlegmariurus crassus B.Øllg.
- Phlegmariurus creber A.R.Field & Bostock
- Phlegmariurus crucis-australis B.Øllg.
- Phlegmariurus cruentus B.Øllg.
- Phlegmariurus cryptomerinus Satou
- Phlegmariurus cuernavacensis B.Øllg.
- Phlegmariurus cumingii B.Øllg.
- Phlegmariurus cuneifolius B.Øllg.
- Phlegmariurus cunninghamioides Ching
- Phlegmariurus curiosus A.R.Field & Bostock
- Phlegmariurus curvifolius B.Øllg.
- Phlegmariurus dacrydioides A.R.Field & Bostock
- Phlegmariurus dalhousieanus A.R.Field & Bostock
- Phlegmariurus darwinianus B.Øllg.
- Phlegmariurus delavayi comb. ined.
- Phlegmariurus delbrueckii A.R.Field & Bostock
- Phlegmariurus deminuens B.Øllg.
- Phlegmariurus dentatus Arana
- Phlegmariurus dianae B.Øllg.
- Phlegmariurus dichaeoides B.Øllg.
- Phlegmariurus dichotomus W.H.Wagner
- Phlegmariurus dielsii A.R.Field & Bostock
- Phlegmariurus divergens A.R.Field & Testo
- Phlegmariurus durus A.R.Field & Bostock
- Phlegmariurus echinatus B.Øllg.
- Phlegmariurus ellenbeckii A.R.Field & Bostock
- Phlegmariurus elmeri A.R.Field & Bostock
- Phlegmariurus engleri B.Øllg.
- Phlegmariurus eremorum B.Øllg.
- Phlegmariurus ericifolius B.Øllg.
- Phlegmariurus erosus comb. ined.
- Phlegmariurus erythrocaulon comb. ined.
- Phlegmariurus espinosanus B.Øllg.
- Phlegmariurus eversus B.Øllg.
- Phlegmariurus fargesii Ching
- Phlegmariurus filicaulon A.R.Field & Testo
- Phlegmariurus filiformis W.H.Wagner
- Phlegmariurus firmus B.Øllg.
- Phlegmariurus flagellaceus A.R.Field & Bostock
- Phlegmariurus flexibilis B.Øllg.
- Phlegmariurus foliaceus B.Øllg.
- Phlegmariurus foliosus A.R.Field & Bostock
- Phlegmariurus fontinaloides B.Øllg.
- Phlegmariurus fordii Ching
- Phlegmariurus friburgensis B.Øllg.
- Phlegmariurus funiculus comb. ined.
- Phlegmariurus funiformis B.Øllg.
- Phlegmariurus gagnepainianus A.R.Field & Bostock
- Phlegmariurus galapagensis B.Øllg.
- Phlegmariurus giganteus A.R.Field & Bostock
- Phlegmariurus gnidioides A.R.Field & Bostock
- Phlegmariurus goebelii A.R.Field & Bostock
- Phlegmariurus goudotii B.Øllg.
- Phlegmariurus guandongensis Ching
- Phlegmariurus gunturensis A.R.Field & Bostock
- Phlegmariurus haeckelii A.R.Field & Testo
- Phlegmariurus haitensis comb. ined.
- Phlegmariurus hamiltonii Á.Löve & D.Löve
- Phlegmariurus harmsii A.R.Field & Bostock
- Phlegmariurus hartwegianus B.Øllg.
- Phlegmariurus hastatus B.Øllg.
- Phlegmariurus hellwigii A.R.Field & Bostock
- Phlegmariurus hemleri B.Øllg.
- Phlegmariurus henryi Ching
- Phlegmariurus heterocarpus B.Øllg.
- Phlegmariurus heteroclitus B.Øllg.
- Phlegmariurus hexastichus B.Øllg.
- Phlegmariurus hildebrandtii A.R.Field & Bauret
- Phlegmariurus hippurideus B.Øllg.
- Phlegmariurus hippuris A.R.Field & Testo
- Phlegmariurus hoffmannii B.Øllg.
- Phlegmariurus hohenackeri B.Øllg.
- Phlegmariurus holstii A.R.Field & Bostock
- Phlegmariurus homocarpus B.Øllg.
- Phlegmariurus horizontalis A.R.Field & Bostock
- Phlegmariurus huberi B.Øllg.
- Phlegmariurus hugonis B.Øllg.
- Phlegmariurus humbertii A.R.Field & Bostock
- Phlegmariurus humbertii-henrici A.R.Field & Bostock
- Phlegmariurus hypogaeus B.Øllg.
- Phlegmariurus hystrix B.Øllg.
- Phlegmariurus ignambiensis A.R.Field & Testo
- Phlegmariurus iminii Kiew
- Phlegmariurus innocentius comb. ined.
- Phlegmariurus insularis comb. ined.
- Phlegmariurus intermedius B.Øllg.
- Phlegmariurus itambensis B.Øllg.
- Phlegmariurus jaegeri A.R.Field & Bostock
- Phlegmariurus kajewskii A.R.Field & Testo
- Phlegmariurus keysseri comb. ined.
- Phlegmariurus killipii B.Øllg.
- Phlegmariurus × koolauensis W.H.Wagner
- Phlegmariurus kuesteri B.Øllg.
- Phlegmariurus lancifolius B.Øllg.
- Phlegmariurus lauterbachii A.R.Field & Bostock
- Phlegmariurus lecomteanus A.R.Field & Bostock
- Phlegmariurus ledermannii A.R.Field & Bostock
- Phlegmariurus lehnertii Testo
- Phlegmariurus leptodon comb. ined.
- Phlegmariurus lignosus B.Øllg.
- Phlegmariurus lindenii B.Øllg.
- Phlegmariurus linifolius B.Øllg.
- Phlegmariurus llanganatensis B.Øllg.
- Phlegmariurus lockyeri A.R.Field & Bostock
- Phlegmariurus loefgrenianus B.Øllg.
- Phlegmariurus longus A.R.Field & Bostock
- Phlegmariurus loxensis B.Øllg.
- Phlegmariurus luederwaldtii comb. ined.
- Phlegmariurus luteynii B.Øllg.
- Phlegmariurus macbridei B.Øllg.
- Phlegmariurus macgregorii A.R.Field & Bostock
- Phlegmariurus magnusianus A.R.Field & Testo
- Phlegmariurus mandiocanus B.Øllg.
- Phlegmariurus mannii W.H.Wagner
- Phlegmariurus marsupiiformis A.R.Field & Bostock
- Phlegmariurus martii B.Øllg.
- Phlegmariurus megastachyus A.R.Field & Bostock
- Phlegmariurus melanesicus A.R.Field & Testo
- Phlegmariurus merrillii A.R.Field & Bostock
- Phlegmariurus mesoamericanus B.Øllg.
- Phlegmariurus mexicanus B.Øllg.
- Phlegmariurus mildbraedii A.R.Field & Bostock
- Phlegmariurus mingjoui X.C.Zhang
- Phlegmariurus minutifolius A.R.Field & Bostock
- Phlegmariurus mirabilis A.R.Field & Testo
- Phlegmariurus mollicomus B.Øllg.
- Phlegmariurus molongensis B.Øllg.
- Phlegmariurus monticola Kiew
- Phlegmariurus mooreanus B.Øllg.
- Phlegmariurus multifarius A.R.Field & Bostock
- Phlegmariurus myrsinites B.Øllg.
- Phlegmariurus myrtifolius A.R.Field & Bostock
- Phlegmariurus myrtuosus B.Øllg.
- Phlegmariurus neocaledonicus A.R.Field & Bostock
- Phlegmariurus nesselii B.Øllg.
- Phlegmariurus niligaricus A.R.Field & Bostock
- Phlegmariurus nudus B.Øllg.
- Phlegmariurus nummularifolius Ching
- Phlegmariurus nutans W.H.Wagner
- Phlegmariurus nyalamensis H.S.Kung & L.B.Zhang
- Phlegmariurus obovalifolius A.R.Field & Testo
- Phlegmariurus obtusifolius A.R.Field & Bostock
- Phlegmariurus ocananus B.Øllg.
- Phlegmariurus oceanianus A.R.Field & Bostock
- Phlegmariurus oellgaardii B.Øllg.
- Phlegmariurus oltmannsii A.R.Field & Bostock
- Phlegmariurus ophioglossoides A.R.Field & Bostock
- Phlegmariurus orizabae B.Øllg.
- Phlegmariurus ovatifolius W.M.Chu ex H.S.Kung & Li Bing Zhang
- Phlegmariurus pachyphyllus A.R.Field & Testo
- Phlegmariurus pachyskelos B.Øllg.
- Phlegmariurus parksii A.R.Field & Bostock
- Phlegmariurus patentissimus A.R.Field & Bostock
- Phlegmariurus pearcei B.Øllg.
- Phlegmariurus pecten A.R.Field & Bostock
- Phlegmariurus perrierianus A.R.Field & Bostock
- Phlegmariurus pflanzii B.Øllg.
- Phlegmariurus phlegmaria Holub
- Phlegmariurus phlegmarioides A.R.Field & Bostock
- Phlegmariurus phylicifolius B.Øllg.
- Phlegmariurus phyllanthus R.D.Dixit
- Phlegmariurus picardae B.Øllg.
- Phlegmariurus pichianus A.R.Field & Bostock
- Phlegmariurus pinifolius Kiew
- Phlegmariurus pithyoides B.Øllg.
- Phlegmariurus pittieri B.Øllg.
- Phlegmariurus podocarpensis B.Øllg.
- Phlegmariurus polaris comb. ined.
- Phlegmariurus polycarpos B.Øllg.
- Phlegmariurus polycladus comb. ined.
- Phlegmariurus polydactylus B.Øllg.
- Phlegmariurus polylepidetorum B.Øllg.
- Phlegmariurus pringlei B.Øllg.
- Phlegmariurus proliferus A.R.Field & Bostock
- Phlegmariurus pruinosus B.Øllg.
- Phlegmariurus pseudovaria comb. ined.
- Phlegmariurus pulcherrimus Á. & D.Löve
- Phlegmariurus pungentifolius B.Øllg.
- Phlegmariurus quadrifariatus B.Øllg.
- Phlegmariurus recurvifolius B.Øllg.
- Phlegmariurus reflexus B.Øllg.
- Phlegmariurus regnellii B.Øllg.
- Phlegmariurus ribourtii A.R.Field & Bostock
- Phlegmariurus riobambensis B.Øllg.
- Phlegmariurus robustus B.Øllg.
- Phlegmariurus rosenstockianus B.Øllg.
- Phlegmariurus rostrifolius B.Øllg.
- Phlegmariurus ruber B.Øllg.
- Phlegmariurus rubricus A.R.Field & Bostock
- Phlegmariurus rufescens B.Øllg.
- Phlegmariurus rupicola A.R.Field & Bostock
- Phlegmariurus sagasteguianus B.Øllg.
- Phlegmariurus salvinioides Ching
- Phlegmariurus samoanus A.R.Field & Bostock
- Phlegmariurus sarmentosus B.Øllg.
- Phlegmariurus saururus B.Øllg.
- Phlegmariurus scabridus B.Øllg.
- Phlegmariurus schlechteri A.R.Field & Bostock
- Phlegmariurus schlimii B.Øllg.
- Phlegmariurus schmidtchenii B.Øllg.
- Phlegmariurus sellifolius B.Øllg.
- Phlegmariurus sellowianus B.Øllg.
- Phlegmariurus serpentiformis B.Øllg.
- Phlegmariurus setifolius A.R.Field & Bostock
- Phlegmariurus shangsiensis C.Y.Yang
- Phlegmariurus sieberianus B.Øllg.
- Phlegmariurus sieboldii Ching
- Phlegmariurus silveirae B.Øllg.
- Phlegmariurus silverstonei B.Øllg.
- Phlegmariurus sintenisii B.Øllg.
- Phlegmariurus societensis A.R.Field
- Phlegmariurus socratis comb. ined.
- Phlegmariurus sooianus Lawalrée
- Phlegmariurus sotae B.Øllg.
- Phlegmariurus sphagnicola B.Øllg.
- Phlegmariurus squarrosus Á. & D.Löve
- Phlegmariurus staudtii A.R.Field & Bostock
- Phlegmariurus stemmermanniae A.C.Medeiros & Wagner
- Phlegmariurus stephani B.Øllg.
- Phlegmariurus strictus A.R.Field & Bostock
- Phlegmariurus subfalciformis A.R.Field & Bostock
- Phlegmariurus subintegrus comb. ined.
- Phlegmariurus subtrifoliatus A.R.Field & Bostock
- Phlegmariurus subtubulosus comb. ined.
- Phlegmariurus subulatus B.Øllg.
- Phlegmariurus tafanensis comb. ined.
- Phlegmariurus talamancanus B.Øllg.
- Phlegmariurus talamauanus A.R.Field & Bostock
- Phlegmariurus talpiphilus B.Øllg.
- Phlegmariurus tardieuae A.R.Field & Testo
- Phlegmariurus tauri A.R.Field & Testo
- Phlegmariurus taxifolius Á. & D.Löve
- Phlegmariurus tenuicaulis B.Øllg.
- Phlegmariurus tenuis B.Øllg.
- Phlegmariurus teretirigidus comb. ined.
- Phlegmariurus terrae-guilelmii A.R.Field & Bostock
- Phlegmariurus tetragonus B.Øllg.
- Phlegmariurus tetrastichoides A.R.Field & Bostock
- Phlegmariurus tetrastichus A.R.Field & Bostock
- Phlegmariurus toppingii A.R.Field & Bostock
- Phlegmariurus tournayanus Lawalrée
- Phlegmariurus transilla B.Øllg.
- Phlegmariurus treitubensis B.Øllg.
- Phlegmariurus trifoliatus A.R.Field & Bostock
- Phlegmariurus trigonus A.R.Field & Bostock
- Phlegmariurus tryonorum B.Øllg.
- Phlegmariurus tubulosus B.Øllg.
- Phlegmariurus ulei comb. ined.
- Phlegmariurus ulixis B.Øllg.
- Phlegmariurus unguiculatus B.Øllg.
- Phlegmariurus urbani B.Øllg.
- Phlegmariurus vanuatuensis A.R.Field
- Phlegmariurus varius A.R.Field & Bostock
- Phlegmariurus venezuelanicus B.Øllg.
- Phlegmariurus vernicosus Á. & D.Löve
- Phlegmariurus villonacensis B.Øllg.
- Phlegmariurus warneckei A.R.Field & Testo
- Phlegmariurus watsonianus B.Øllg.
- Phlegmariurus weberbaueri B.Øllg.
- Phlegmariurus weddellii B.Øllg.
- Phlegmariurus wilsonii B.Øllg.
- Phlegmariurus xiphophyllus A.R.Field & Bostock
- Phlegmariurus yunnanensis Ching