Haplogroup O-M119
In human genetics, Haplogroup O-M119 is a Y-chromosome DNA haplogroup. Haplogroup O-M119 is a descendant branch of haplogroup O-F265, one of two extant primary subclades of Haplogroup O-M175. The same clade previously has been labeled as O-MSY2.2.
Origins
The Haplogroup O-M119 branch is believed to have evolved during the Late Pleistocene in China mainland.Paleolithic migrations
A 2010 study by Karafet suggests haplogroup O-M119 was part of a four-phase colonization model in which Paleolithic migrations of hunter-gatherers shaped the primary structure of current Y-Chromosome diversity of Maritime Southeast Asia. Neolithic incursions made only a minor impact on the paternal gene pool, despite the large cultural impact of the Austronesian expansion. Approximately 5000 BCE, Haplogroup O-M119 coalesced at Sundaland and migrated northwards to as far as Taiwan, where O-M50 constitutes some 90% of the Aboriginal Y-DNA, being the main haplogroup that can be directly linked to the Austronesian expansion in phase 3.Taiwan homeland
A study by Li in 2008 concluded that in contrast to the Taiwan homeland hypothesis, Island Southeast Asians do not have a Taiwan origin based on their paternal lineages. According to their results, lineages within Maritime Southeast Asia did not originate from Taiwanese aborigines as linguistic studies suggest. Taiwan aborigines and Indonesians were likely to have been derived from the Tai–Kadai-speaking populations based on their paternal lineages, and thereafter evolved independently of each other.The strongest positive correlation between Haplogroup O-M119 and ethno-linguistic affiliation is that which is observed between this haplogroup and the Austronesians. The peak frequency of Haplogroup O-M119 is found among the aborigines of Taiwan, precisely the region from which linguists have hypothesized that the Austronesian language family originated. A slightly weaker correlation is observed between Haplogroup O-M119 and the Han Chinese populations of southern China, as well as between this haplogroup and the Tai–Kadai-speaking populations of southern China and Southeast Asia. The distribution of Tai–Kadai languages in Thailand and other parts of Southeast Asia outside of China has long been believed, for reasons of traditional linguistic geography, to reflect a recent invasion of Southeast Asia by Tai–Kadai-speaking populations originating from southeastern China, and the somewhat elevated frequency of Haplogroup O-M119 among the Tai–Kadai populations, coupled with a high frequency of Haplogroup O-M95, which is a genetic characteristic of the Austroasiatic-speaking peoples of Southeast Asia, suggests that the genetic signature of the Tai–Kadai peoples' affinity with populations of southeastern China has been weakened due to extensive assimilation of the earlier Austroasiatic residents of the lands which the Tai–Kadai peoples invaded. Also, it has been noted that Haplogroup O-M119 lineages among populations of continental Southeast Asia outside of China display a reduced level of diversity when compared with populations of South China and insular Southeast Asia, which may be evidence of a bottleneck associated with the westward migration and settlement of ancestral Tai–Kadai-speaking populations in Indochina.
Distribution
Haplogroup O-M119 lineages are found primarily in Southeast Asian populations of Malaysia, Vietnam, Indonesia, the Philippines, southern China and Taiwan. High frequencies of this haplogroup have been found in populations spread in an arc through southeastern China, Taiwan, the Philippines, and Indonesia. It has been found with generally lower frequency in samples from Oceania, mainland Southeast Asia, Southwest China, Northwest China, North China, Northeast China, Korea, Japan, North Asia, and Central Asia.Population | Percentage | Count | Source | SNPs |
Nias | 100.0% | 60 | P203=52 M110=8 | |
Nias | 99.8% | 407 | M119 | |
Taiwanese aborigines | 89.6% | 48 | P203=34 M110=9 | |
Mentawai | 86.5% | 74 | M119=42 P203=21 M110=1 | |
Aboriginal Taiwanese | 83.4% | 223 | M119 | |
Taiwan | 78.0% | 41 | M119=18 M110=14 | |
Taiwan | 71.8% | 39 | M119=15 M50=13 | |
Taiwan | 68.9% | 74 | M119 | |
Atayal | 62.5% | 24 | M119=13 M50/M110/M103=2 | |
Utsat | 61.1% | 72 | M119=44 | |
Gelao | 60.0% | 30 | M119=18 | |
Gelong | 57.7% | 78 | M119=45 | |
Tagalog | 46.0% | 50 | M119 | |
Kota Kinabalu | 42.1% | 19 | M119=6 M50/M110/M103=2 | |
Philippines | 41.0% | 39 | M119=11 M110=5 | |
Hlai | 40.0% | 50 | M119=20 | |
Philippines | 35.7% | 28 | M119=9 M50=1 | |
Dong | 35% | 20 | M119=5 M110/M50/M103=2 | |
Hlai | 32.0% | 75 | M119=24 | |
Sui | 31.5% | 92 | M119=29 | |
Malaysian | 30.8% | 13 | M110=3 M119=1 | |
Dong | 30% | 10 | M119=2 M50/M110/M103=1 | |
Kota Kinabalu | 29.2% | 65 | M119=12 M50=6 M101=1 | |
Hlai | 28.8% | 66 | M119=19 | |
Trobriand Islands | 28.3% | 53 | M119=15 | |
Hlai | 27.3% | 11 | M119=3 | |
Trobriand Islands | 26.9% | 52 | M110=9 M119=5 | |
Li | 26.5% | 34 | M119 | |
Malay | 25.0% | 12 | M119 | |
Han | 24.0% | 167 | M119 | |
Han Chinese | 23.1% | 26 | M119=6 | |
Hlai | 23.0% | 74 | M119=17 | |
Banjarmasin | 22.7% | 22 | M119=3 M50=2 | |
Java | 22.6% | 53 | M119=10 M110=2 | |
Nusa Tenggara | 22.6% | 31 | M119=4 M110=3 | |
Batak | 22.2% | 18 | M119=4 | |
China | 22.2% | 36 | M119=8 | |
Balinese | 21.1% | 641 | P203=73 M119=57 M110=5 | |
Mandar | 20.4% | 54 | M119=7 M110=4 | |
Tujia | 20% | - | - | |
Han | 20.0% | 35 | M119 | |
Buka | 20.0% | 10 | M119 | |
CDX | 19.2% | 52 | K644/Z23266=7 F656=2 Z23442=1 | |
Zhuang | 19.0% | 63 | M119=12 | |
Malay | 18.5% | 27 | M50/M110/M103=4 M119=1 | |
Thin Board Mien | 18.2% | 11 | M119 | |
Majuro | 18.2% | 11 | M50=2 | |
Balinese | 18.1% | 551 | M119=100 | |
Sui | 18.0% | 50 | M119=9 | |
Zhuang | 17.9% | 28 | M119=5 | |
Admiralty Islands | 17.7% | 147 | M110=26 | |
Malaysia | 17.6% | 17 | M119=3 | |
Sumatra | 17.5% | 57 | M119=10 | |
Malagasy | 17.1% | 35 | M50 | |
Han | 16.9% | 65 | M119 | |
Qiang | 15.2% | 33 | M119 | |
Borneo | 15.0% | 40 | M119=5 M110=1 | |
Han Chinese | 15% | - | - | |
Dai | 15.0% | 20 | M119=3 | |
Java | 14.8% | 61 | P203=6 M119=2 M110=1 | |
She | 14.7% | 34 | M119 | |
Han | 14.7% | 34 | M119 | |
Manus | 14.3% | 7 | M119 | |
Palyu | 13.3% | 30 | M119 | |
Batak Toba | 13.2% | 38 | M119=4 P203=1 | |
Sumba | 12.6% | 350 | M110=22 M119=19 P203=3 | |
Micronesia | 12.5% | 16 | M119=2 | |
Guizhou Miao | 12.2% | 49 | M119 | |
Lowland Kimmun | 12.2% | 41 | M119 | |
Thai | 11.8% | 17 | P203 | |
Tai Yong | 11.5% | 26 | P203=2 M50=1 | |
Bunu | 11.1% | 36 | M119 | |
Malaysia | 11.1% | 18 | M119=2 | |
Filipinos | 10.4% | 48 | M110=4 P203=1 | |
Zhuang | 10% | - | - | |
Mountain Straggler Mien | 10.0% | 20 | M119 | |
Northeast Thai | 10.0% | 20 | M119=1 M50/M110/M103=1 | |
Vietnam | 10% | 10 | M119=1 | |
Tai Lue | 9.9% | 91 | P203=6 M50=3 | |
Han | 9.7% | 165 | P203=15 M119=1 | |
Flores | 9.6% | 394 | P203=36 M119=2 | |
Zhuang | 9.6% | 166 | - | |
Han | 9.5% | 21 | M119 | |
Han | 9.4% | 32 | M119 | |
Borneo | 9.3% | 86 | M110=5 P203=2 M119=1 | |
Bougainville | 9.2% | 65 | M119 | |
Top Board Mien | 9.1% | 11 | M119 | |
Northern Mien | 9.1% | 33 | M119 | |
Northern She | 8.9% | 56 | M119 | |
Thai | 8.8% | 34 | M119 | |
Hui | 8.6% | 35 | M119 | |
Mosuo | 8.5% | 47 | M119 | |
Miao | 8.3% | 48 | M119=4 | |
Tonga | 8.3% | 12 | M119=1 | |
Tujia | 8.2% | 49 | P203=4 | |
Hlai | 8.1% | 62 | M119=5 | |
Hlai/Cun | 8% | - | - | |
Cambodian | 7.7% | 26 | M119=1 M50/M110/M103=1 | |
Ewenki | 7.7% | 26 | M119 | |
Xibe | 7.3% | 41 | M119 | |
Dai | 7.1% | 28 | M119=2 | |
Hunan Miao | 7.0% | 100 | M119 | |
Tujia | 7% | - | - | |
Miao | 6.9% | 58 | P203=4 | |
Bai | 6.7% | 30 | M119=2 | |
Katu | 6.7% | 45 | M119 | |
Moluccas | 6.7% | 30 | P203=1 M110=1 | |
Han | 6.7% | 30 | M119 | |
Kinh | 6.7% | 15 | M119 | |
Kinh | 6.6% | 76 | P203 | |
Southern Mien | 6.5% | 31 | M119 | |
Yao | 6.4% | 47 | M119=3 | |
Malaysia | 6.3% | 32 | M119=1 P203=1 | |
Dai | 6.1% | 49 | M119=3 | |
Lisu | 6.1% | 49 | M119=3 | |
Yunnan Miao | 6.1% | 49 | M119 | |
Northern Han | 6.1% | 49 | M119 | |
Comorians | 6.0% | 381 | M50=22 MSY2.2=1 | |
Bai | 6.0% | 50 | M119 | |
Bai | 6.0% | 50 | M119=3 | |
Moluccas | 5.9% | 34 | M119=1 M110=1 | |
Kyrgyz | 5.8% | 52 | M119 | |
Vietnam | 5.7% | 70 | P203=4 | |
Yao | 5.7% | 35 | M119 | |
Fiji | 5.6% | 107 | M119=6 | |
Mon | 5.6% | 18 | P203=1 | |
Samoa | 5.6% | 18 | P203=1 | |
Thailand | 5.3% | 75 | P203=2 M119=2 | |
Daur | 5.1% | 39 | M119 | |
Cham | 5.1% | 59 | M119 | |
Dungan | 5.0% | 40 | M119 | |
Shan | 5.0% | 20 | P203=1 | |
Manchu | 4.9% | 41 | M119=2 | |
Han | 4.8% | 42 | M119 | |
Maewo | 4.5% | 44 | M119=1 M110=1 | |
Bouyei | 4.4% | 45 | M119=2 | |
Jino | 4.4% | 45 | M119=2 | |
Korean | 4.4% | 45 | M119 | |
KHV | 4.3% | 46 | Z23392=1 Z23442=1 | |
Han | 4.3% | 47 | M119=2 | |
Western Mien | 4.3% | 47 | M119 | |
Pumi | 4.3% | 47 | M119 | |
Zhuang | 4.3% | 47 | M119=2 | |
Buyi | 4.2% | 48 | M119=2 | |
Mongolian | 4.2% | 24 | M119 | |
Tai Khün | 4.2% | 24 | P203=1 | |
Korea | 4.0% | 25 | M119=1 | |
Western Samoa | 4.0% | 25 | M119=1 | |
Khon Mueang | 3.9% | 205 | P203=6 M50=2 | |
Manchu | 3.8% | 52 | M119 | |
Koreans | 3.8% | 133 | P203=3 M119=2 | |
New Ireland | 3.7% | 109 | M119 | |
Bo | 3.6% | 28 | M119 | |
Tai Yuan | 3.5% | 85 | P203=3 | |
Japanese | 3.4% | 263 | M119 | |
Lembata | 3.3% | 92 | M119=2 P203=1 | |
Korean | 3.2% | 216 | M119 | |
Samoa | 3.2% | 62 | M119=2 | |
Han | 3.1% | 129 | M119 | |
Papua New Guinea | 3.0% | 33 | P203=1 | |
Manchu | 3.0% | 101 | M119 | |
Koreans | 3.0% | 573 | P203=16 M119=1 | |
Dai | 2.9% | 35 | M119=1 | |
Manchu | 2.9% | 35 | M119 | |
Han | 2.9% | 35 | M119 | |
Buyi | 2.9% | 35 | M119 | |
Yao | 2.9% | 35 | M119 | |
West New Britain | 2.8% | 249 | M119 | |
Koreans | 2.7% | 300 | M119 | |
Koreans | 2.7% | 75 | M119 | |
Japanese | 2.6% | 117 | M119 | |
Koreans | 2.4% | 85 | M119 | |
Lavongai | 2.3% | 43 | M119 | |
Koreans | 2.2% | 506 | M119 | |
Laven | 2.0% | 50 | M119 | |
Yi | 2.0% | 50 | M119 | |
Hmong Daw | 2.0% | 51 | M119 | |
She | 2.0% | 51 | P203=1 | |
Japanese | 1.9% | 104 | M119 | |
Vanuatu | 1.9% | 52 | M50=1 | |
Yao | 1.7% | 60 | P203=1 | |
Uygur | 1.4% | 70 | M119 | |
East New Britain | 1.4% | 145 | M119 | |
Japanese | 1.2% | 2390 | M119 | |
Tuvalu | 1.0% | 100 | M110=1 | |
Mongolia | 0.7% | 149 | M119 | |
Mongols | 0.6% | 160 | M119 | |
Lawa | 0.0% | 50 | M119=0 |
A 2008 study by Li suggested that the admixture analyses of Tai–Kadai-speaking populations showed a significant genetic influence in a large proportion of Indonesians. Most of the population samples contained a high frequency of haplogroup O-M119.
The frequencies of Haplogroup O-M119 among various East Asian and Austronesian populations suggest a complex genetic history of the modern Han populations of southern China. Although Haplogroup O-M119 occurs only at an average frequency of approximately 4% among Han populations of northern China and peoples of southwestern China and Southeast Asia who speak Tibeto-Burman languages, the frequency of this haplogroup among the Han populations of southern China nearly quadruples to about 15-23%. The frequency of Haplogroup O-M119 among the Southern Han has been found to be slightly greater than the arithmetic mean of the frequencies of Haplogroup O-M119 among the Northern Han and a pooled sample of Austronesian populations. This suggests that modern Southern Han populations may possess a non-trivial number of male ancestors who were originally affiliated with some Austronesian-related culture, or who at least shared some genetic affinity with many of the ancestors of modern Austronesian peoples.
Subclade distribution
O-M119
This lineage is found frequently in Austronesians, southern Han Chinese, and Kra-Dai peoples. This lineage is presumed to be a marker of the prehistoric Austronesian expansion, with possible origins encompassing the regions along the southeastern coast of China and neighboring Taiwan, and is found among modern populations of Maritime Southeast Asia and Oceania.Haplogroup O-M119 Y-chromosomes also have been found to occur at low frequency among various populations of Siberia, such as the Nivkhs, Ulchi/Nanai, Yenisey Evenks, and especially the Buryats living in the Sayan-Baikal uplands of Irkutsk Oblast .
O-P203
O-P203 was found in 86.7% of a sample from Nias, 70.8% of Taiwanese Aboriginals, 28.4% of Mentawai, 11.4% of Balinese, 9.8% of a sample from Java, 9.1% of a sample from Flores, 9.1% of Han Chinese, 8.3% of a sample from Western Samoa, 8.2% of Tujia from Hunan, 6.9% of Miao from China, 5.7% of Vietnamese, 3.3% of a sample from the Moluccas, 3.1% of Malaysians, 3.0% of a sample from highland Papua New Guinea, 2.6% of a sample from Sumatra, 2.3% of a sample from Borneo, 2.1% of Filipinos, 2.0% of She, 1.7% of Yao from Guangxi, 1.1% of a sample from Lembata, and 0.9% of a sample from Sumba.In a study published in 2011, O-P203 was observed in 22.2% of Han Chinese male volunteers at Fudan University in
Shanghai whose origin may be traced back to East China, 12.3% of Han Chinese male volunteers whose origin may be traced back to South China, and 1.6% of Han Chinese male volunteers whose origin may be traced back to North China.
O-M101
This lineage was observed in one individual from China and another from Kota Kinabalu.O-M50
This lineage occurs among Austronesian peoples of Taiwan, the Philippines, Indonesia, Melanesia, Micronesia, and Madagascar as well as among some populations of continental Southeast Asia and among Bantu peoples of the Comoros. It also has been found in a Hawaiian.A study published in 2005 found O-M50 in 33.3% of a sample of aboriginals in Taiwan, 18.2% of a sample of people in Majuro, 17.1% of a sample of Malagasy, 9.2% of a sample of people in Kota Kinabalu, 9.1% of a sample of people in Banjarmasin, 3.6% of a sample of people in the Philippines, and 1.9% of a sample of people in Vanuatu.
Kayser et al. 2008 found O-M110 in 34.1% of a sample of Taiwan Aborigines, 17.7% of a sample from the Admiralty Islands, 17.3% of a sample from the Trobriand Islands, 13.5% of a sample from the Philippines, 9.7% of a sample from the Nusa Tenggara Islands, 3.8% of a sample from Java, 3.0% of a sample from the Moluccas, 2.5% of a sample from Borneo, 1.0% of a sample from Tuvalu, and 0.95% of a sample from Fiji.
A study published in 2010 found O-M110 in 18.8% Taiwanese Aboriginals, 13.3% Nias, 8.3% Philippines, 7.4% Sulawesi, 6.3% Sumba, 5.8% Borneo, 3.3% Moluccas, 2.3% Maewo, Vanuatu, 1.6% Java, 1.4% Mentawai, and 0.8% Bali.
A study published in 2012 found O-M110 in 4.6% of males from the Solomon Islands.
Phylogenetics
Phylogenetic history
Prior to 2002, there were in academic literature at least seven naming systems for the Y-Chromosome Phylogenetic tree. This led to considerable confusion. In 2002, the major research groups came together and formed the Y-Chromosome Consortium. They published a joint paper that created a single new tree that all agreed to use. Later, a group of citizen scientists with an interest in population genetics and genetic genealogy formed a working group to create an amateur tree aiming at being above all timely. The table below brings together all of these works at the point of the landmark 2002 YCC Tree. This allows a researcher reviewing older published literature to quickly move between nomenclatures.YCC 2002/2008 | ' | ' | ' | ' | ' | ' | YCC 2002 | YCC 2005 | YCC 2008 | YCC 2010r | ISOGG 2006 | ISOGG 2007 | ISOGG 2008 | ISOGG 2009 | ISOGG 2010 | ISOGG 2011 | ISOGG 2012 | |
O-M175 | 26 | VII | 1U | 28 | Eu16 | H9 | I | O* | O | O | O | O | O | O | O | O | O | O |
O-M119 | 26 | VII | 1U | 32 | Eu16 | H9 | H | O1* | O1a | O1a | O1a | O1a | O1a | O1a | O1a | O1a | O1a | O1a |
O-M101 | 26 | VII | 1U | 32 | Eu16 | H9 | H | O1a | O1a1 | O1a1a | O1a1a | O1a1 | O1a1 | O1a1a | O1a1a | O1a1a | O1a1a | O1a1a |
O-M50 | 26 | VII | 1U | 32 | Eu16 | H10 | H | O1b | O1a2 | O1a2 | O1a2 | O1a2 | O1a2 | O1a2 | O1a2 | O1a2 | O1a2 | O1a2 |
O-P31 | 26 | VII | 1U | 33 | Eu16 | H5 | I | O2* | O2 | O2 | O2 | O2 | O2 | O2 | O2 | O2 | O2 | O2 |
O-M95 | 26 | VII | 1U | 34 | Eu16 | H11 | G | O2a* | O2a | O2a | O2a | O2a | O2a | O2a | O2a | O2a | O2a1 | O2a1 |
O-M88 | 26 | VII | 1U | 34 | Eu16 | H12 | G | O2a1 | O2a1 | O2a1 | O2a1 | O2a1 | O2a1 | O2a1 | O2a1 | O2a1 | O2a1a | O2a1a |
O-SRY465 | 20 | VII | 1U | 35 | Eu16 | H5 | I | O2b* | O2b | O2b | O2b | O2b | O2b | O2b | O2b | O2b | O2b | O2b |
O-47z | 5 | VII | 1U | 26 | Eu16 | H5 | I | O2b1 | O2b1a | O2b1 | O2b1 | O2b1a | O2b1a | O2b1 | O2b1 | O2b1 | O2b1 | O2b1 |
O-M122 | 26 | VII | 1U | 29 | Eu16 | H6 | L | O3* | O3 | O3 | O3 | O3 | O3 | O3 | O3 | O3 | O3 | O3 |
O-M121 | 26 | VII | 1U | 29 | Eu16 | H6 | L | O3a | O3a | O3a1 | O3a1 | O3a1 | O3a1 | O3a1 | O3a1 | O3a1 | O3a1a | O3a1a |
O-M164 | 26 | VII | 1U | 29 | Eu16 | H6 | L | O3b | O3b | O3a2 | O3a2 | O3a2 | O3a2 | O3a2 | O3a2 | O3a2 | O3a1b | O3a1b |
O-M159 | 13 | VII | 1U | 31 | Eu16 | H6 | L | O3c | O3c | O3a3a | O3a3a | O3a3 | O3a3 | O3a3a | O3a3a | O3a3a | O3a3a | O3a3a |
O-M7 | 26 | VII | 1U | 29 | Eu16 | H7 | L | O3d* | O3c | O3a3b | O3a3b | O3a4 | O3a4 | O3a3b | O3a3b | O3a3b | O3a2b | O3a2b |
O-M113 | 26 | VII | 1U | 29 | Eu16 | H7 | L | O3d1 | O3c1 | O3a3b1 | O3a3b1 | - | O3a4a | O3a3b1 | O3a3b1 | O3a3b1 | O3a2b1 | O3a2b1 |
O-M134 | 26 | VII | 1U | 30 | Eu16 | H8 | L | O3e* | O3d | O3a3c | O3a3c | O3a5 | O3a5 | O3a3c | O3a3c | O3a3c | O3a2c1 | O3a2c1 |
O-M117 | 26 | VII | 1U | 30 | Eu16 | H8 | L | O3e1* | O3d1 | O3a3c1 | O3a3c1 | O3a5a | O3a5a | O3a3c1 | O3a3c1 | O3a3c1 | O3a2c1a | O3a2c1a |
O-M162 | 26 | VII | 1U | 30 | Eu16 | H8 | L | O3e1a | O3d1a | O3a3c1a | O3a3c1a | O3a5a1 | O3a5a1 | O3a3c1a | O3a3c1a | O3a3c1a | O3a2c1a1 | O3a2c1a1 |
Research publications
The following research teams per their publications were represented in the creation of the YCC tree.Phylogenetic trees
This phylogenetic tree of haplogroup O subclades is based on the YCC 2008 tree and subsequent published research.- O-MSY2.2
- *O-M119
- **O1a-Y144065
- ***O1a-Y144065* Hubei
- ***O1a-F1009 Guangdong, Gansu, North Korea
- **O1a1
- ***O1a1*
- ***O1a1a
- ****O1a1a1
- *****O1a1a1a
- ******O1a1a1a* Indonesia
- ******O1a1a1a1
- *******O1a1a1a1* China, Philippines
- *******O1a1a1a1a China, Philippines, Vietnam
- ********O1a1a1a1a* Guangdong
- ********O1a1a1a1a1
- *********O1a1a1a1a1a
- **********O1a1a1a1a1a1
- ***********O1a1a1a1a1a1a Hubei, Jiangsu, Hong Kong, Beijing, Xishuangbanna
- ************O1a1a1a1a1a1a1 Jiangsu, Anhui
- *************O1a1a1a1a1a1a1a
- ************O1a1a1a1a1a1a2
- ************O1a1a1a1a1a1a3
- ************O1a1a1a1a1a1a4
- ************O1a1a1a1a1a1a5
- ***********O1a1a1a1a1a1b Guangdong, Guangxi, Jiangsu
- ************O1a1a1a1a1a1b1
- ************O1a1a1a1a1a1b2 Fujian
- ************O1a1a1a1a1a1b3 Hubei, Jiangxi
- ***********O1a1a1a1a1a1c
- ************O1a1a1a1a1a1c* Jiangxi, Fujian
- ************O1a1a1a1a1a1c1 Sichuan, Cambodia
- ***********O1a1a1a1a1a1d Henan, Anhui
- ***********O1a1a1a1a1a1e Jiangsu, Anhui
- ************O1a1a1a1a1a1e1
- **********O1a1a1a1a1a2 Jiangsu, Beijing
- ********O1a1a1a1a2 Jiangsu, Shanghai
- ******O1a1a1a2
- *******O1a1a1a2a
- ********O1a1a1a2a1 Philippines, Singapore
- *********O1a1a1a2a1a Philippines, Singapore
- *******O1a1a1a2b
- *****O1a1a1b
- ******O1a1a1b1 China, Kota Kinabalu
- ******O1a1a1b2
- *******O1a1a1b2* Ho Chi Minh City
- *******O1a1a1b2a Xishuangbanna, Hong Kong, Ho Chi Minh City
- ********O1a1a1b2a1
- ****O1a1a2
- *****O1a1a2a Japan
- ******O1a1a2a1
- *******O1a1a2a1a
- ********O1a1a2a1a* Fujian
- ********O1a1a2a1a1 Guangdong, Sichuan, Hunan, Beijing
- ********O1a1a2a1a2 Xishuangbanna
- ********O1a1a2a1a3 Xishuangbanna
- *******O1a1a2a1b
- ********O1a1a2a1b* Shandong
- ********O1a1a2a1b1 Beijing, Guangdong, Liaoning
- ******O1a1a2a2
- *******O1a1a2a2a Zhejiang, Shandong
- *******O1a1a2a2b Hubei, Jiangxi, Zhejiang, Hebei, North Korea
- *****O1a1a2b
- ******O1a1a2b1 Jiangsu, Shanghai
- ******O1a1a2b2 Jiangsu, Shanghai, Hungary
- ***O1a1b Beijing, Fujian, the Philippines, Singapore
- **O1a2 Austronesia, China
- ***O1a2a
- ****O1a2a1
- *****O1a2a1a
Genetics
Y-DNA O subclades
Y-DNA backbone tree
Footnotes
Works cited
JournalsWebsites