It was described as a new species in 1825 by Dutch zoologist Coenraad Jacob Temminck. Temminck acquired the specimens used for his description from Dutch businessman Jan Cock Blomhoff Its species name "dasymallus" is likely ; Temminck described its fur as long and woolly. The five subspecies are:
Daito fruit bat - P. d. daitoensis
Erabu fruit bat - P. d. dasymallus
Taiwanese fruit bat - P. d. formosus
Orii's fruit bat - P. d. inopinatus
Yaeyama fruit bat - P. d. yayeyamae
The subspecies are based on populations that occur on different islands.
Description
The Ryukyu flying fox is slightly smaller than the Indian flying fox, with a wingspan of. It weighs. Its forearm is approximately long. The body of the bat is covered in long hairs, making the body seem almost woolly. The bat is reddish brown and has a yellowish white nape. Its ears are small and pointed, and are difficult to see beneath its thick fur. Its flight membranes are dark brown in color.
Biology and ecology
It is mostly frugivorous, consuming the fruits of at least 53 plant species; the flowers of 20 plant species; the leaves of 18 plant species; and the bark of one plant species. It has also been observed consuming eight different species of insect. The Chinesebanyan tree is an important source of food year-round. It is an important pollinator of a subspecies of Schima wallichii, an evergreen tree. It also pollinates a species of climbing vine, Mucuna macrocarpa. It is a nocturnal species, usually solitary roosting in trees during the day and foraging at night. The Ryukyu flying fox enhances seed dispersal, as seeds from digested fruits are deposited as guano up to from the parent trees.
The largest population of these bats is probably on the Philippines and is thought to be stable. In Japan there are estimated to be well over five thousand individuals but in Taiwan, there has been a large reduction in bat numbers. This species faces a number of threats. Some populations in the Philippines are hunted for consumption and this bat is considered a delicacy on Babuyan Claro. In Japan, habitat loss is the main threat but some individuals get entangled in nets placed to protect citrus crops and others are electrocuted by power-lines. Overall, most populations have been in decline though this seemed to have levelled off to some extent by 2008 when the IUCN removed this bat from the "Endangered" category and placed it in the "Vulnerable" category.
Relationship to humans
In Temminck's initial description, he wrote that it "devastates" orchards. Its depredation on orchards caused Okinawa Prefecture to launch an investigation in 2012. In two villages surveyed in 2013, it was estimated that flying foxes cause 19 million yen in damages to citrus crops annually. Many Japanese farmers believe that the Ryukyu flying fox is a pest that should be managed by culling.