Porcelain crab Porcelain crabs are decapod crustaceans in the widespread family Porcellanidae , which superficially resemble true crabs. They have flattened bodies as an adaptation for living in rock crevices. They are delicate, readily losing limbs when attacked, and use their large claws for maintaining territories. They first appeared in the Tithonian age of the Late Jurassic epoch , 145-152 million years ago .Description Porcelain crabs are small, usually with body widths less than. They share the general body plan of a squat lobster , but their bodies are more compact and flattened, an adaptation for living and hiding under rocks. Porcelain crabs are quite fragile animals, and often shed their limbs to escape predators, hence their name. The lost appendage can grow back over several moults . Porcelain crabs have large chelae , which are used for territorial struggles, but not for catching food. The fifth pair of pereiopods is reduced and used for cleaning.Evolution Porcelain crabs are an example of carcinisation , whereby a noncrab-like animal evolves into an animal that resembles a true crab . Porcelain crabs can be distinguished from true crabs by the apparent number of walking legs , and the long antennae originating on the front outside of the eyestalks. The abdomen of the porcelain crab is long and folded underneath it, free to move.Biogeography and ecology Porcelain crabs live in all the world's oceans , except the Arctic Ocean and the Antarctic . They are common under rocks, and can often be found and observed on rocky beaches and shorelines, startled creatures scurrying away when a stone is lifted. They feed by combing plankton and other organic particles from the water using long setae on the mouthparts . Some of the common species of porcelain crabs in the Caribbean Sea are Petrolisthes quadratus , found in large numbers under rocks in the intertidal , and the red-and-white polka-dotted Porcellana sayana , which lives commensally within the shells inhabited by large hermit crabs . In Hong Kong , Petrolisthes japonicus is common.Diversity , some 4723 extant species porcelain crab had been described, divided among these 30 genera:Aliaporcellana Nakasone & Miyake, 1969Allopetrolisthes Haig , 1960Ancylocheles Haig, 1978Capilliporcellana Haig, 1978Clastotoechus Haig, 1960Enosteoides Johnson, 1970Euceramus Stimpson, 1860Eulenaios Ng & Nakasone, 1993Heteropolyonyx Osawa, 2001Heteroporcellana Haig, 1978Liopetrolisthes Haig, 1960Lissoporcellana Haig, 1978Madarateuchus Harvey, 1999Megalobrachium Stimpson, 1858 Minyocerus Stimpson, 1858Neopetrolisthes Miyake, 1937Neopisosoma Haig, 1960Novorostrum Osawa, 1998Orthochela Glassell, 1936Pachycheles Stimpson, 1858Parapetrolisthes Haig, 1962Petrocheles Miers, 1876Petrolisthes Stimpson, 1858Pisidia Leach, 1820Polyonyx Stimpson, 1858Porcellana Lamarck, 1801Porcellanella White, 1852Pseudoporcellanella Sankarankutty, 1962Raphidopus Stimpson, 1858Ulloaia Glassell, 1938 The fossil record of porcelain crabs includes species of Pachycheles , Pisidia , Polyonyx , Porcellana , and a further six genera known only from fossils:Annieporcellana Fraaije et al. , 2008Beripetrolisthes De Angeli & Garassino, 2002Eopetrolisthes De Angeli & Garassino, 2002Jurellana Schweitzer & Feldmann, 2010Lobipetrolisthes De Angeli & Garassino, 2002Longoporcellana Müller & Collins , 1991 The earliest porcelain crab fossil is Jurellana from the Tithonian of central Europe.
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