Piksi


Piksi is a potential pterosaur genus containing the single species Piksi barbarulna. It lived roughly 75 million years ago in what is now Montana, United States. Known from parts of a right wing – the humerus, ulna and radius bones – the only specimens found so far are housed in the Museum of the Rockies. The genus Piksi is monotypic at present.
The fossils were found in 1991 by Gloria Jean Siebrecht in the Blackfeet Indian Reservation, namely at Bob's Vacation Site locality TM-088, Glacier County. Recovered from an old stratum of the upper Two Medicine Formation, they are probably from an individual that died in or near a small pool. It was described in 2002 by David J. Varricchio.

Description and classification

The bones are fragmentary and represent roughly the elbow area. Comparing the fossils' size to the wing bones of other ground birds, P. barbarulna seems to have been about as large as a common pheasant, i.e. some 35–40 cm long excluding tail, and with a wingspan of perhaps 80 cm or somewhat less. It would thus have weighed some 500 g – 1 kg.
The original description of the fossils found its affinities unresolvable except that it was probably an ornithothoracine bird. Agnolin and Varricchio reinterpreted Piksi barbarulna as a pterosaur rather than a bird, most likely a member of Ornithocheiroidea. However, it has since been noted that the humerus of Piksi possesses features which found in some theropods but not in any pterodactyloid pterosaurs.
However, more recent studies have recovered it as a pterosaur again.

Ecology

The deposit in which the bones were found was a silty claystone. This was formed from sediments deposited during what seems to have been a rather cool phase of the Late Cretaceous: sea levels of the Western Interior Seaway at least were apparently very low for Mesozoic standards, though this may also have been due to strong tectonic uplift in the Cordilleran Overthrust Belt. The location was inland, with the Western Interior Seaway's coast at least 350 km away.
Judging from the stratigraphical, sedimentological and faunal data, the environment was thus probably semi-humid, possibly semi-arid grassland or shrubland in a tropical or subtropical climate. The claystone apparently formed from sediment of a small floodplain, such as an ephemeral pool.
A diverse fauna utilized the location as a habitat. Other theropods were plentiful, including Troodon, tyrannosaurids, and dromaeosaurids. Orodromeus was also abundant; herds might have come to the pool to drink or breed, as adults, juveniles and hatchlings were found together. Early mammals – metatherians and multituberculates – occurred in the area, as well as lizards. That there was a temporary though not permanent body of water is indicated by the presence of articulated frog skeletons and the absence of fish and other aquatic animals.