Oystercatcher


The oystercatchers are a group of waders forming the family Haematopodidae, which has a single genus, Haematopus. They are found on coasts worldwide apart from the polar regions and some tropical regions of Africa and South East Asia. The exceptions to this are the Eurasian oystercatcher, the South Island oystercatcher, and the Magellanic oystercatcher, which also breed inland, far inland in some cases. In the past there has been a great deal of confusion as to the species limits, with discrete populations of all black oystercatchers being afforded specific status but pied oystercatchers being considered one single species.
The name oystercatcher was coined by Mark Catesby in 1731 as a common name for the North American species H. palliatus, described as eating oysters. Yarrell in 1843 established this as the preferred term, replacing the older name sea pie or sea-pie. The genus name Haematopus comes from the Greek haima blood, pous foot.

Description

The different species of oystercatcher show little variation in shape or appearance. They range from in length and in wingspan. The Eurasian oystercatcher is the lightest on average, at, while the sooty oystercatcher is the heaviest, at. The plumage of all species is either all-black, or black on top and white underneath. The variable oystercatcher is slightly exceptional in being either all-black or pied. They are large, obvious, and noisy plover-like birds, with massive long orange or red bills used for smashing or prying open molluscs. The bill shape varies between species, according to the diet. Those birds with blade-like bill tips pry open or smash mollusc shells, and those with pointed bill tips tend to probe for annelid worms. They show sexual dimorphism, with females being longer-billed and heavier than males.

Feeding

The diet of oystercatchers varies with location. Species occurring inland feed upon earthworms and insect larvae. The diet of coastal oystercatchers is more varied, although dependent upon coast type; on estuaries, bivalves, gastropods and polychaete worms are the most important part of the diet, whereas rocky shore oystercatchers prey upon limpets, mussels, gastropods, and chitons. Other prey items include echinoderms, fish, and crabs.

Breeding

Nearly all species of oystercatcher are monogamous, although there are reports of polygamy in the Eurasian oystercatcher. They are territorial during the breeding season. There is strong mate and site fidelity in the species that have been studied, with one record of a pair defending the same site for 20 years. A single nesting attempt is made per breeding season, which is timed over the summer months. The nests of oystercatchers are simple affairs, scrapes in the ground which may be lined, and placed in a spot with good visibility. The eggs of oystercatchers are spotted and cryptic. Between one and four eggs are laid, with three being typical in the Northern Hemisphere and two in the south. Incubation is shared but not proportionally, females tend to take more incubation and males engage in more territory defence. Incubation varies by species, lasting between 24–39 days. Oystercatchers are also known to practice "egg dumping." Like the cuckoo, they sometimes lay their eggs in the nests of other species such as seagulls, abandoning them to be raised by those birds.

Conservation

The Canary Islands oystercatcher became extinct during the 20th century. The Chatham oystercatcher is endemic to the Chatham Islands of New Zealand but is listed as endangered by the IUCN, while both the African and Eurasian oystercatchers are considered near threatened. There has been conflict with commercial shellfish farmers, but studies have found that the impact of oystercatchers is much smaller than that of shore crabs.

Species

Several fossil species are known, including Haematopus sulcatus from the early Pliocene of Florida. The oldest known fossil species is Haematopus palliatus, known from the Barstovian of Florida and Zanclean of North Carolina.