Nerve tract


A nerve tract is a bundle of nerve fibers connecting nuclei of the central nervous system. In the peripheral nervous system this is known as a nerve, and has associated connective tissue. The main nerve tracts in the central nervous system are of three types: association fibers, commissural fibers, and projection fibers. A tract may also be referred to as a commissure, fasciculus or decussation. A commissure connects the two cerebral hemispheres at the same levels. Examples are the posterior commissure and the corpus callosum. A decussation is a connection made by fibres that cross at different levels, such as the sensory decussation. Examples of a fascicle are the subthalamic fasciculus and the lenticular fasciculus.

Types

The nerve fibers in the central nervous system can be categorized into three groups on the basis of their course and connections. Different tracts may also be referred to as projections or radiations.

Association fibers

The tracts that connect cortical areas within the same hemisphere are called association tracts. Long association fibers connect different lobes of a hemisphere to each other whereas short association fibers connect different gyri within a single lobe. Among their roles, association tracts link perceptual and memory centers of the brain.
The cingulum is a major association tract. The cingulum forms the white matter core of the cingulate gyrus and links from this to the entorhinal cortex. Another major association tract is the superior longitudinal fasciculus that has three parts.

Commissural fibers

connect corresponding cortical areas in the two hemispheres. They cross from one cerebral hemisphere to the other through bridges called commissures. The great majority of commissural tracts pass through the largest commissure the corpus callosum. A few tracts pass through the much smaller anterior and posterior commissures. Commissural tracts enable the left and right sides of the cerebrum to communicate with each other. Other commissures are the hippocampal commissure, and the habenular commissure.

Projection fibers

connect the cerebral cortex with the corpus striatum, diencephalon, brainstem and the spinal cord. The corticospinal tract for example, carries motor signals from the cerebrum to the spinal cord. Other projection tracts carry signals upward to the cerebral cortex. Superior to the brainstem, such tracts form a broad, dense sheet called the internal capsule between the thalamus and basal nuclei, then radiate in a diverging, fanlike array to specific areas of the cortex.