Mouse-eared bat
The mouse-eared bats are a diverse and widespread genus of bats within the family Vespertilionidae.
Relationships
Myotis has historically been included in the subfamily Vespertilioninae, but was classified in its own subfamily, Myotinae, by Nancy Simmons in 1998. In her 2005 classification in Mammal Species of the World, Simmons listed the genera Cistugo and Lasionycteris in the Myotinae in addition to Myotis itself. However, molecular data indicate that Cistugo is distantly related to all other Vespertilionidae, so it was reclassified into its own family, the Cistugidae, and that Lasionycteris belongs in the Vespertilioninae. The genus Submyotodon has since been added to the subfamily, making it and Myotis its only members.Appearance and behavior
Their ears are normally longer than they are wide, with a long and lance-shaped tragus, hence their English and zoological names. The species within this genus vary in size from very large to very small for vesper bats, with a single pair of mammary glands.Mouse-eared bats are generally insectivores. M. vivesi, and several members of the trawling bat ecomorph Leuconoe, have relatively large feet with long toes, and take small fish from the water surface.
Longevity
Myotis species are remarkably long-lived for their size; in 2018, researchers revealed that a longitudinal study appears to indicate that Myotis telomeres do not shrink with age, and that telomerase does not appear to be present in the Myotis metabolism.Species
Traditionally, Myotis has been divided into three large subgenera—Leuconoe, Myotis, and Selysius. However, molecular data indicate that these subgenera are not natural groups, but instead unnatural assemblages of convergently similar species. Instead, Myotis species largely fall in two main clades, one containing Old World and the other New World species. However, the Asian species Myotis latirostris falls outside the clade formed by these main groups, and may represent a separate genus, and the Eurasian Myotis brandtii is related to New World species.Myotis is a highly species-rich genus, and the classification of many species remains unsettled. In the below list, all differences in taxonomy from the 2005 third edition of Mammal Species of the World are indicated in footnotes.
- Myotis adversus - large-footed bat, large-footed mouse-eared bat, large-footed myotis
- Myotis aelleni - southern myotis
- Myotis albescens - silver-tipped myotis
- Myotis alcathoe - Alcathoe bat
- Myotis altarium - Szechwan myotis
- Myotis anjouanensis - Anjouan myotis
- Myotis annamiticus - Annamit myotis
- Myotis annectans - hairy-faced bat
- Myotis atacamensis - Atacama myotis
- Myotis ater - Peters's myotis, small black myotis
- Myotis attenboroughi Moratelli et al., 2017 - Sir David Attenborough's myotis
- Myotis aurascens
- Myotis auriculus - southwestern myotis
- Myotis australis - Australian myotis
- Myotis austroriparius - southeastern myotis
- Myotis bechsteinii - Bechstein's bat
- Myotis blythii - lesser mouse-eared bat
- Myotis bocagii - rufous mouse-eared bat
- Myotis bombinus - Far Eastern myotis, bombinus bat
- Myotis brandtii - Brandt's bat
- Myotis bucharensis - Bocharic myotis, Bokhara whiskered bat
- Myotis californicus - California myotis
- Myotis capaccinii - long-fingered bat
- Myotis chiloensis - Chilean myotis
- Myotis chinensis - large myotis
- Myotis ciliolabrum - western small-footed bat, western small-footed myotis
- Myotis cobanensis - Guatemalan myotis
- Myotis crypticus Ruedi, Ibáñez, Salicini, Juste & Puechmaille, 2019 - cryptic myotis
- Myotis csorbai - Csorba's mouse-eared bat
- Myotis dasycneme - pond bat
- Myotis daubentonii - Daubenton's bat
- Myotis davidii - David's myotis
- Myotis dieteri - Kock's mouse-eared bat
- Myotis diminutus Moratelli & Wilson, 2011
- Myotis dominicensis Miller, 1902 - Dominican myotis
- Myotis elegans Hall, 1962 - elegant myotis
- Myotis emarginatus - Geoffroy's bat
- Myotis escalerai Cabrera, 1904 - Escalera's bat
- Myotis evotis - long-eared myotis
- Myotis fimbriatus - fringed long-footed myotis
- Myotis findleyi Bogan, 1978 - Findley's myotis
- Myotis flavus Shamel, 1944
- Myotis formosus - Hodgson's bat, copper-winged bat
- Myotis fortidens Miller and Allen, 1928 - cinnamon myotis
- Myotis frater G.M. Allen, 1923 - fraternal myotis
- Myotis gomantongensis Francis and Hill, 1998 - Gomantong myotis
- Myotis goudoti - Malagasy mouse-eared bat
- Myotis gracilis Ognev, 1927
- Myotis grisescens A.H. Howell, 1909 - gray bat
- Myotis hajastanicus Argyropulo, 1939 - Armenian whiskered bat, Hajastan myotis, Armenian myotis
- Myotis hasseltii - lesser large-footed bat
- Myotis hermani Thomas, 1923 - Herman's myotis
- Myotis horsfieldii - Horsfield's bat
- Myotis ikonnikovi Ognev, 1912 - Ikonnikov's bat
- Myotis insularum - insular myotis
- Myotis izecksohni Moratelli, Peracchi, Dias & de Oliveira, 2011
- Myotis keaysi J.A. Allen, 1914 - hairy-legged myotis
- Myotis keenii - Keen's myotis
- Myotis laniger Peters, 1871 - Chinese water myotis
- Myotis leibii - eastern small-footed bat
- Myotis levis - yellowish myotis
- Myotis longipes - Kashmir cave bat
- Myotis lucifugus - little brown bat
- Myotis macrodactylus - eastern long-fingered bat, big-footed myotis
- Myotis macropus - southern myotis, large-footed myotis
- Myotis macrotarsus - pallid large-footed myotis, Philippine large-footed myotis
- Myotis martiniquensis LaVal, 1973 - Schwartz's myotis
- Myotis melanorhinus Merriam, 1890 - dark-nosed small-footed myotis
- Myotis midastactus Moratelli & Wilson, 2014
- Myotis moluccarum - Maluku myotis, Arafura large-footed bat
- Myotis montivagus - Burmese whiskered bat
- Myotis morrisi Hill, 1971 - Morris's bat
- Myotis muricola - wall-roosting mouse-eared bat, Nepalese whiskered myotis
- Myotis myotis - greater mouse-eared bat
- Myotis mystacinus - whiskered bat
- Myotis nattereri - Natterer's bat
- Myotis nesopolus Miller, 1900 - Curacao myotis
- Myotis nigricans - black myotis
- Myotis nipalensis Dobson, 1871 - Nepal myotis
- Myotis nyctor LaVal & Schwartz, 1974
- Myotis occultus Hollister, 1909 - Arizona myotis
- Myotis oreias - Singapore whiskered bat
- Myotis oxyotus - montane myotis
- Myotis peninsularis Miller, 1898 - peninsular myotis
- Myotis pequinius Thomas, 1908 - Beijing mouse-eared bat, Peking myotis
- Myotis petax Hollister, 1912 - eastern water bat, Sakhalin bat
- Myotis phanluongi Borisenko, Kruskop and Ivanova, 2008
- Myotis pilosus Peters, 1869 - Rickett's big-footed bat
- Myotis planiceps Baker, 1955 - flat-headed myotis
- Myotis pruinosus Yoshiyuki, 1971 - frosted myotis
- Myotis punicus Felten, Spitzenberger and Storch, 1977 - Felten's myotis
- Myotis ricketti
- Myotis ridleyi Thomas, 1898 - Ridley's bat
- Myotis riparius Handley, 1960 - riparian myotis
- Myotis rosseti - thick-thumbed myotis
- Myotis ruber - red myotis
- Myotis rufoniger
- Myotis schaubi Kormos, 1934 - Schaub's myotis
- Myotis scotti Thomas, 1927 - Scott's mouse-eared bat
- Myotis septentrionalis - northern long-eared bat, northern myotis
- Myotis sicarius Thomas, 1915 - Mandelli's mouse-eared bat
- Myotis siligorensis - Himalayan whiskered bat
- Myotis simus Thomas, 1901 - velvety myotis
- Myotis sodalis Miller and Allen, 1928 - Indiana bat
- Myotis stalkeri Thomas, 1910 - Kei myotis
- Myotis taiwanensis Ärnbäck-Christie-Linde, 1908
- Myotis thysanodes Miller, 1897 - fringed myotis
- Myotis tricolor - Cape hairy bat, little brown bat, Temminck's mouse-eared bat, Cape myotis, tricoloured mouse-eared bat, Cape hairy myotis, Temminck's hairy bat, three-coloured bat
- Myotis velifer - cave myotis
- Myotis vivesi Menegaux, 1901 - fish-eating bat, fish-eating myotis
- Myotis volans - long-legged myotis
- Myotis welwitschii - Welwitsch's bat, Welwitsch's mouse-eared bat, Welwitsch's myotis
- Myotis yanbarensis Maeda and Matsumara, 1998 - Yanbaru whiskered bat
- Myotis yumanensis - Yuma myotis
- Myotis zenatius Ibáñez, Juste, Salicini, Puechmaille & Ruedi, 2019 -Zenati myotis
Literature cited
- Borisenko, A.V., Kruskop, S.V. and Ivanova, N.V. 2008. A new mouse-eared bat from Vietnam. Russian Journal of Theriology 7:57–69.
- Han, N., Zhang, J., Reardon, T., Lin, L., Zhang, J. and Zhang, S. 2010. . Acta Chiropterologica 12:449–456.*Happold, M. 2005. A new species of Myotis from central Africa. Acta Chiropterologica 7:9–21.
- Ibáñez, C., García-Mudarra, J.L., Ruedi, M., Stadelmann, B. and Juste, J. 2006. The Iberian contribution to cryptic diversity in European bats. Acta Chiropterologica 8:277–297.
- Jiang, T., Sun, K., Chou, C., Zhang, Z. and Feng, J. 2010. First record of Myotis flavus from mainland China and a reassessment of its taxonomic status. Zootaxa 2414:41–51.
- Lack, J.B., Roehrs, Z.P., Stanley, C.E., Ruedi, M. and Van Den Bussche, R.A. 2010. . Journal of Mammalogy 91:976–992.
- Matveev, V.A., Kruskop, S.V. and Kramerov, D.A. 2005. Revalidation of Myotis petax Hollister, 1912 and its new status in connection with M. daubentonii . Acta Chiropterologica 7:23–37.
- Mayer, F., Dietz, C. and Kiefer, A. 2007. . Frontiers in Zoology 4:239–255.
- Roehrs, Z.P., Lack, J.B. and Van Den Bussche, R.A. 2010. . Journal of Mammalogy 91:1073–1092.
- Simmons, N.B. 2005. Order Chiroptera. Pp. 312–529 in Wilson, D.E. and Reeder, D.M.. . 3rd ed. Baltimore: The Johns Hopkins University Press, 2 vols., 2142 pp.
- Stadelmann, B., Lin, L.-K., Kunz, T.H. and Ruedi, M. 2007. . Molecular Phylogenetics and Evolution 43:32–48.
- Tsytsulina, K. 2004. On the taxonomical status of Myotis abei Yoshikura, 1944. Zoological Science 21:963–966.