The Middleton Formation is the eastern correlate of the Teekloof Formation. Both these formations overlie the Abrahamskraal Formation. The upper Middleton Formation correlates with eastern exposures of the uppermost PristerognathusAssemblage Zone, and is late Middle Permian in age. Current thought posits that the eruption of the Emeishan Large Igneous Province caused the middlePermian extinction event, during which the Middleton Formation rocks formed. The Middleton Formation, like the underlying Abrahamskraal Formation, is rich in mudstone. However, the mudstones are notably redder in colour in the Middleton formation than in the Abrahamskraal Formation. The Middleton mudstones contain wave-and current-ripples, as well as calcareousnodules, which weather out brown. Minor sandstones, containing argillaceous layers, interspace the mudstones. These sandstone layers are important stratigraphic markers for geologists and paleontologists. The mudstones were deposited in a shallow, low-energy, freshwater lacustrine environment. Observation of the sandstones reveals that high-energy fluvial events infrequently disturbed this low-energy depositional environment. These disturbances are associated with extensive argillaceous overbank deposits. The argillaceous sandstone deposits contain many trace fossils of invertebrates, suggesting that the deposits were nutrient-rich.
Paleontology
The Middleton Formation does not contain as many fossils as its western correlate, the Teekloof Formation. This is particularly true of vertebrate fossils. The end-Guadalupian extinction event caused a drop in species diversity in the Middleton Formation compared to the Abrahamskraal Formation. However, vertebrate fossils yielded from the Middleton Formation are the same as those associated with the Pristerognathus Assemblage Zone. When found, vertebrate fossils appear most frequently in the mudstone and calcareous nodule deposits. Trace fossils are more commonly found in this formation than vertebrate fossils. These trace fossils lie in the sandstones and include preserved burrows, track ways, feeding trails and other evidence of shallow-water-dwelling invertebrates. The invertebrates include annelids, aquatic oligochaetes, nematodes, insect larvae, and planolites. The formation's trace fossils also offer evidence for the presence of the bony fishesNamaichthys and Atherstonia.