In the late 1990s, Osiel Cárdenas Guillén, the former leader of the Gulf cartel, had other similar groups besides Los Zetas established in several cities in Tamaulipas. Each of these groups were identified by their radio codes: the Rojos were based in Reynosa; the Metros were headquartered in Matamoros; and the Lobos were established in Laredo. The infighting between the Metros and the Rojos of the Gulf cartel began in 2010, when Juan Mejía González, nicknamed El R-1, was overlooked as the candidate of the regional boss of Reynosa and was sent to the "Frontera Chica," an area that encompasses Miguel Alemán, Camargo and Ciudad Mier – directly across the U.S.–Mexico border from Starr County, Texas. The area that Mejía González wanted was given to Samuel Flores Borrego, suggesting that the Metros were above the Rojos. Unconfirmed information released by The Monitor indicated that two leaders of the Rojos, Mejía González and Rafael Cárdenas Vela, teamed up to kill Flores Borrego. Cárdenas Vela had held a grudge on Flores Borrego and the Metros because he believed that they had led the Mexican military to track down and kill his uncle Antonio Cárdenas Guillén on 5 November 2010. Other sources indicate that the infighting could have been caused by the suspicions that the Rojos were "too soft" on the Gulf cartel's bitter enemy, Los Zetas. When the Gulf cartel and Los Zetas split in early 2010, some members of the Rojos stayed with the Gulf cartel, while others decided to leave and join the forces of Los Zetas. InSight Crime explains that the fundamental disagreement between the Rojos and the Metros was over leadership. Those who were more loyal to the Cárdenas family stayed with the Rojos, while those loyal to Jorge Eduardo Costilla Sánchez, like Flores Borrego, defended the Metros. Originally, the Gulf cartel was running smoothly, but the infighting between the two factions in the Gulf cartel triggered when Flores Borrego was killed on 2 September 2011. When the Rojos turned on the Metros, the largest faction in the Gulf cartel, firefights broke throughout Tamaulipas and drug loads were stolen among each other, but the Metros managed to retain control of the major cities that stretched from Matamoros to Miguel Alemán, Tamaulipas.