List of fossil bird genera

evolved from certain feathered theropod dinosaurs, and there is no real dividing line between birds and non-avian dinosaurs except that some of the former survived the Cretaceous–Paleogene extinction event while the latter did not. For the purposes of this article, a 'bird' is considered to be any member of the clade Aves in the broadest sense. Some dinosaur groups which may or may not be true birds are listed below under Proto-birds.
This page contains a listing of prehistoric bird taxa only known from completely fossilized specimens. These extinctions took place before the Late Quaternary and thus took place in the absence of significant human interference. While the earliest hominids had been eating birds and especially their eggs, human population and technology was simply insufficient to seriously affect healthy bird populations until the Upper Paleolithic Revolution. Rather, reasons for the extinctions listed here are stochastic abiotic events such as bolide impacts, climate changes due to orbital shifts, mass volcanic eruptions etc. Alternatively, species may have gone extinct due to evolutionary displacement by successor or competitor taxa – it is notable that an extremely large number of seabirds have gone extinct during the mid-Tertiary; this seems at least partly due to competition by the contemporary radiation of marine mammals.
The relationships of these taxa are often hard to determine, as many are known only from very fragmentary remains and due to the complete fossilization precluding analysis of information from DNA, RNA or protein sequencing. The taxa listed in this article should be classified with the "Fossil".
Before the late 19th century, when minerals were still considered one of the kingdoms of binomial nomenclature, fossils were often treated according to a parallel taxonomy. Rather than assigning them to animal or plant genera, they were treated as mineral genera and given binomial names typically using Osteornis or Ornitholithus as "genus". The latter name, however, is still in use for an oogenus of fossil bird eggs. Also, other animals were placed in these "genera". In sources pre-dating the Linnean system, the above terms are also seen in the more extensive descriptions used to name taxa back then.

Taxonomic list of fossil prehistoric birds

Higher-level taxa are presented in likely or suspected phylogenetic order. Genus-level taxa and lower are sorted chronologically, in ascending order.
The higher-level groups of non-Neornithes are arranged based on the phylogeny proposed by Luis Chiappe, updated and expanded to incorporate recent research. The categories are inclusive in ascending order.
Taxonomic assignments, especially in the pygostylian to early neornithine genera, are still very provisional and subject to quite frequent change.

Basal [Avialae] (extinct)

The most primitive "birds", usually still possessing a long bony tail with generally unfused vertebrae. Not all of these may be on the line of bird ancestors; whether they are not closer to other theropods groups than to the Avialae remains to be thoroughly tested.

†Anchiornis huxleyi Xu et al. 2009
†Aurornis xui Godefroit et al. 2013
†Balaur bondoc Csiki et al. 2010
†Jixiangornis orientalis Ji et al. 2002
†Xiaotingia zhengi Xu et al. 2011
†Yandangornithiformes Cai & Zhao 1999
* †Yandangornithidae Cai & Zhao 1999
** †Yandangornis longicaudatus Cai & Zhao 1999
†Archaeopterygiformes Fürbringer 1888
* †Archaeopterygidae Huxley 1871 a
** †Wellnhoferia grandis Elzanowski 2001 – possible synonym of Archaeopteryx
** †Archaeopteryx Meyer 1861
†Jeholornithiformes Zhou & Zhang 2006
* †Jeholornithidae Zhou & Zhang 2006
** †Dalianraptor cuhe Gao & Liu 2005
** †Jeholornis Zhou & Zhang 2002b
Basal [Pygostylia] (extinct)

The earliest birds with a modern pygostyle: a reduction and fusion of the tail vertebrae; possibly a paraphyletic group. Two types of pygostyle are known, a rod-shaped one found in Confuciusornithidae, Enantiornithes and some non-avian theropods such as Nomingia, and a plowshare-shaped one, only known in the lineage leading to modern birds. It is not certain that the pygostyles found in birds are indeed synapomorphies.

†Chongmingia zhengi Wang et al. 2016
†Confuciusornithiformes Hou et al. 1995
* †Proornis coreae Lim 1993
* †Zhongornis haoae Gao et al. 2008 - confuciusornithid?
* †Confuciusornithidae Hou et al. 1995
** †Changchengornis hengdaoziensis Ji, Chiappe & Ji 1999
** †Eoconfuciusornis zhengi Zhang, Zhou & Benton 2008
** †Evgenavis nobilis O’Connor, Averianov & Zelenkov 2014
** †Confuciusornis Hou et al. 1995
†Omnivoropterygiformes Czerkas & Ji 2002
* †Omnivoropterygidae Czerkas & Ji 2002
** †Omnivoropteryx sinousaorum Czerkas & Qiang Ji 2002
** †Sapeornis chaoyangensis Zhou & Zhang 2002a
[Enantiornithes] (extinct)

The taxonomic list of enantiornithine groups presented here follows a summary published by Thomas R. Holz, Jr. in 2011.
†Enantiornithes

Basal Enantiornithes and Enantiornithes incerta sedis
*†Cerebavis cenomanica Kurochkin et al. 2006
*†Cruralispennia multidonta Wang et al. 2017
*†Dalingheornis liweii Zhang et al. 2006
*†Dunhuangia cuii Wang et al. 2015
*†Elsornis keni Chiappe et al. 2007
*†Feitianius paradisi O'Connor et al. 2015
*†Holbotia ponomarenkoi Zelenkov & Averianov 2015
*†Houornis caudatus Wang & Liu 2015
*†Laevisoolithidae Mikhailov 1991
**†Laevisoolithus sochavai Mikhailov 1991
**†Subtiliolithus microtuberculatus Mikhailov 1991
**†Tipoolithus achloujensis Garcia et al. 2003
*†Linyiornis amoena Wang et al. 2016
*†Monoenantiornis sihedangia Hu & O’Connor 2016
*†Paraprotopteryx gracilis Zheng, Zhang & Hou 2007
*†Parvavis chuxiongensis Wang, Zhou & Xu 2014
*†Piscivorenantiornis inusitatus Wang & Zhou 2017
*†Pterygornis dapingfangensis Wang, Hu & Li 2015
*†Xiangornis shenmi Hu et al. 2012
*†Yuanjiawaornis viriosus Hu et al. 2015
*†Kurzholiidae Nesov 1992
**†Kuszholia mengi Nesov 1992
*†Iberomesornithiformes Sanz & Bonaparte 1992
**†Iberomesornithidae Sanz & Bonaparte 1992
***†Iberomesornis romerali Sanz & Bonaparte 1992
*†Liaoningornithiformes Hou 1997
**†Liaoningornithidae Hou 1997
***†Eoalulavis hoyasi Sanz et al. 1996
***†Liaoningornis longidigitris Hou 1997
Basal Euenantiornithes
*†Catenoleimus anachoretus Panteleyev 1999
*†Cratoavis cearensis Carvalho et al. 2015
*†Elbretornis bonapartei Walker & Dyke 2009
*†Eocathayornis walkeri Zhou 2002
*†Flexomornis howei Tykoski & Fiorillo 2010
*†Fortunguavis xiaotaizicus Wang, O’Connor & Zhou 2014
*†Grabauornis lingyuanensis Dalsatt, Erickson & Zhou 2014
*†Huoshanornis huji Wang et al. 2010
*†Largirostrornis sexdentoris Hou 1997
*†Lectavis bretincola Chiappe 1993
*†Martinavis Walker, Buffetaut & Dyke 2007
*†Yungavolucris brevipedalis Chiappe 1993
*†Liaoxiornithiformes Hou et al. 2002
**†Liaoxiornithidae Hou & Chen 1999
***†Liaoxiornis delicatus Hou & Chen 1999
*†Pengornithidae Wang et al. 2014
**†Chiappeavis magnapremaxillo O’Connor et al. 2016
**†Eopengornis martini Wang et al. 2014
**†Pengornis houi Zhou, Clarke & Zhang 2008
**†Parapengornis eurycaudatus Hu, O’Connor & Zhou 2015
*†Protopterygiformes Zhou & Zhang 2006
**†Protopterygidae Zhou & Zhang 2006
***†Jibeinia luanhera Hou 2000
***†Protopteryx fengningensis Zhang & Zhou 2000
***†Hebeiornis fengningensis Yan 1999
†Eoenantiornithiformes Hou et al. 1999
*†Eoenantiornithidae Hou et al. 1999
**†Dapingfangornis sentisorhinus Li et al. 2006
**†Eoenantiornis buhleri Hou et al. 1999
*†Bohaiornithidae Wang et al. 2014
**†Sulcavis geeorum O’Connor et al. 2013
**†Shenqiornis mengi Wang et al. 2010
**†Zhouornis hani Zhang et al. 2013
**†Longusunguis kurochkini Wang et al. 2014
**†Bohaiornis guoi Hu et al. 2011
**†Parabohaiornis martini Wang et al. 2014
†Longipterygiformes Zhang et al. 2001
*†Longipterygidae Zhang et al. 2001
**†Boluochia zhengi Zhou 1995
**†Camptodontus yangi Li et al. 2010 non Dejean 1826
**†Longipteryx chaoyangensis Zhang et al. 2001
**†Longirostravis hani Hou et al. 2004
**†Otogornis genghisi Hou 1994
**†Rapaxavis pani Morschhauser et al. 2009
**†Shanweiniao cooperorum O’Connor et al. 2009
**†Shengjingornis yangi Li et al. 2012
†Cathayornithiformes Zhou, Jin & Zhang 1992
*†Alethoalaornithidae Li et al. 2007
**†Alethoalaornis agitornis Li et al. 2007
*†Concornithidae Kuročkin 1996
**†Concornis lacustris Sanz & Buscalioni 1992
**†Qiliania graffini Ji et al. 2011
**†Noguerornis gonzalezi Lasaca-Ruiz 1989
*†Cathyornithidae Zhou, Jin & Zhang 1992
**†Gracilornis jiufotangensis Li & Hou 2011
**†Longchengornis sanyanensis Hou 1997
**†Sinornis santensis Sereno & Rao 1992
**†Cathayornis Zhou, Jin & Zhang 1992
†Enantiornithiformes Chiappe 1992 non Walker 1981, Martin 1983
*†Avisauridae Brett-Surman & Paul 1985
**†Cuspirostrisornis houi Hou 1997
**†Avisaurus Brett-Surman & Paul 1985
**†Bauxitornis mindszentyae
**†Enantiophoenix electrophyla Cau & Arduini 2008
**†Halimornis thompsonae Chiappe, Lamb & Ericson 2002
**†Gettyia Atterholt et al., 2018
**†Intiornis inexpectatus
**†Mirarce Atterholt et al., 2018
**†Mystiornis cyrili Kuročkin et al. 2011
**†Neuquenornis volans Chiappe & Calvo 1994
**†Soroavisaurus australis Chiappe 1993
*†Gobipterygidae Elzanowski 1974
**†Gobipipus reshetovi Kuročkin, Chatterjee & Mikhailov 2013
**†Gobipteryx minuta Elzanowski 1974
**†Vescornis hebeiensis Zhang, Ericson & Zhou 2004
*†Enantiornithidae Walker 1981
**†Enantiornis leali Walker 1981
**†Gurilynia nessovi Kuročkin 1999
**†Wyleyia valdensis Harrison & Walker 1973
**†Nanantius Molnar 1986
*†Alexornithidae Brodkorb 1976
**†Abavornis bonaparti Panteleyev 1998
**†Alexornis antecedens Brodkorb 1976
**†Explorornis Panteleyev 1998
**†Incolornis Panteleyev 1998
**†Kizylkumavis cretacea Nesov 1984
**†Lenesornis maltschevskyi Kuročkin 1996
**†Sazavis prisca Nesov & Jarkov 1989
**†Zhyraornis Nesov 1984

Note that Holtz also included Zhyraornis in his classification of euenantiornithines, though this genus is more often classified as an ornithuran. Holtz also placed Liaoningornis as an ornithuromorph, though more recent studies have placed it as a close relative of Eoalulavis.

Basal [Euornithes] (extinct)

Also called "basal Ornithuromorpha". Essentially modern birds, except many still possess a few primitive features such as teeth or wing claws. These have the plowshare-shaped pygostyle and proper tail fan as seen in most living birds. The taxonomy of this group is confusing; the name "Ornithurae" was first proposed by Ernst Haeckel in 1866 and has been revised in meaning several times since.
The following is a list of primitive euornithian genera and those that cannot be confidently referred to any subgroups, following Holtz.

†Archaeorhynchus spathula Zhou & Zhang 2006
†Changmaornis houi Wang et al. 2013
†Changzuiornis ahgmi Huang et al. 2017
†Gargantuavis philoinos Buffetaut & Le Loeuff 1998
†Hollanda luceria Bell et al. 2009
†Horezmavis eocretaceous Nesov 1983
†Hulsanpes perlei Osmólska 1982
†Iteravis huchzermeyeri Zhou, O'Connor & Wang 2014
†Jianchangornis microdonta Zhou, Zhang & Li 2009
†Jiuquanornis niui Wang et al. 2013
†Juehuaornis zhangi Wang, Wang & Hu 2015
†Palaeopteryx thomsoni Jensen 1981
†Platanavis nana Nesov 1992
†Schizooura lii Zhou, Zhou & O'Connor 2012
†Tingmiatornis arctica Wang et al. 2017
†Vorona berivotrensis Forster et al. 1996
†Xinghaiornis lini Wang et al. 2013
†Yumenornis huangi Wang et al. 2013
†Zhongjianornis yangi Zhou, Zhang & Li 2009
†Patagopterygidae Alvarenga & Bonaparte 1992
*†Alamitornis minutus Agnolin & Martinelli 2009
*†Patagopteryx deferrariisi Alvarenga & Bonaparte 1992
†Hongshanornithidae O’Connor, Gao & Chiappe 2010
*?†Archaeornithura meemannae Wang et al. 2015
*†Hongshanornis longicresta Zhou & Zhang 2005
*†Longicrusavis houi O’Connor, Gao & Chiappe 2010
*†Parahongshanornis chaoyangensis Li, Wang & Hou 2011 – ornithurine?
*†Tianyuornis cheni Zheng et al. 2014
†Chaoyangiiformes Hou 1997
*†Chaoyangiidae Hou 1997
**†Chaoyangia beishanensis Hou & Zhang 1993
†Songlingornithiformes
*†Songlingornithidae Hou 1997
**†Piscivoravis lii Zhou et al. 2013
**†Songlingornis linghensis Hou 1997
**†Yixianornis grabaui Zhou & Zhang 2001
**†Yanornis Zhou & Zhang 2001

Note that Holtz also included the genera Eurolimnornis, Holbotia, Palaeocursornis and Piksi as euornitheans, though they have since been re-identified as pterosaurs.

Basal [Ornithurae] (extinct)

Limenavis patagonica Clarke & Chiappe 2001 – paleognath?
Palintropidae
* Palintropus retusus Brodkorb 1970
†Ambiortiformes Kuročkin 1982
*†Ambiortidae Kuročkin 1982
**†Ambiortus dementjevi Kuročkin 1982
†Apsaraviformes Kuročkin 1982
*†Apsaravidae
**†Apsaravis ukhaana Norell & Clarke 2001
†Gansuiformes Hou & Liu 1984
*†Gansuidae Hou & Liu 1984
**†Gansus Hou & Liu 1984
†Ichthyornithiformes Marsh 1873
*†Apatornithidae Fürbringer 1888
**†Apatornis celer Marsh 1873
**†Guildavis tener Clarke 2004
**†Iaceornis marshi Clarke 2004
*†Ichthyornithidae Marsh 1873
** Ichthyornis Marsh 1872
†Hesperornithiformes Fürbringer 1888
*†Chupkaornis keraorum Tanaka et al. 2017
*†Potamornis skutchi Elzanowski, Paul & Stidham 2000
*†Pasquiaornis Tokaryk, Cumbaa & Storer 1997
*†Enaliornithidae Fürbringer 1888
**†Enaliornis Seeley 1876
*†Baptornithidae AOU 1910
**†Baptornis Marsh 1877
*†Brodavidae Martin, Kuročkin & Tokaryk 2012
**Brodavis Martin, Kuročkin & Tokaryk 2012
*†Judinornithidae Nesov 1983
**†Judinornis nogontsavensis Nesov & Borkin 1983
*†Hesperornithidae Marsh 1880
**†Fumicollis hoffmani Bell & Chiappe 2015
**†Parahesperornis alexi Martin 1980
**†Hesperornis'' Marsh 1872
[Neornithes]

The subclass that contains all modern birds.
Unresolved and basal forms

These modern birds are known from remains that cannot be placed in relation to any one modern group and are neither autapomorphic enough to assign them to own orders. Especially the Late Cretaceous/early Paleogene taxa are probably basal to several modern orders, while later Paleogene taxa often represent extinct lineages outside the modern families.

†Australornis lovei Mayr & Scofield 2014
†Gallornis straeleni Lambrecht 1931
†Ceramornis major Brodkorb 1963 – charadriiform?
†"Presbyornithidae" gen. et sp. indet. - anseriform ?
†Torotix clemensi Brodkorb 1963 – pelecaniform, charadriiform, procellariiform or phoenicopteriform
Neornithes incerta sedis Kurochkin 1995 - phalacrocoracid?
Neornithes incerta sedis AMNH FR 25272 - phalacrocoracid?
Neornithes incerta sedis PVPH 237 - galliform?
Neornithes incerta sedis UCMP 117598
Neornithes incerta sedis UCMP 117599 – anseriform?
†"Lonchodytes" pterygius Brodkorb 1963 – charadriiform?
†Novacaesareala hungerfordi Parris & Hope 2002 – related to Torotix?
†"Palaeotringa" vetus Marsh 1870 - gruiform? anseriform ?
†Volgavis marina Nessov & Jarkow 1989 – charadriiform? phalacrocoraciform?
†Tshulia litorea Nessov 1988
†Eupterornis remensis Lemoine 1878 – charadriiform ? gaviiform?
†Gradiornis walbeckensis Mayr 2007 – cariamid?
†"Messelornis" russelli Mourer-Chauviré 1995 sensu Mayr 2007 – messelornithid?
†Walbeckornis creber Mayr 2007 – charadriiform? messelornithid?
Neornithes incerta sedis – charadriiform, ciconiiform, gruiform?
†Argillipes Harrison & Walker 1977 – galliform?
*†A. aurorum Harrison & Walker 1977
*†A. magnus Harrison & Walker 1979
*†A. paralectoris Harrison & Walker 1977
†Coturnipes cooperi Harrison & Walker 1977 – galliform, falconiform?
†Fluviatilavis antunesi Harrison 1983 – charadriiform?
†Mopsitta tanta Waterhouse et al. 2008 – threskiornithid, psittacid?
†Neanis schucherti Shufeldt 1913 – coraciiform, piciform?
†Paleophasianus meleagroides Wetmore 1940 – galliform or gruiform ?
†Precursor parvus – several species? psittaciform + charadriiform ?
†"Precursor" litorum Harrison & Walker 1977
†"Precursor" magnus Harrison & Walker 1977
†Procuculus minutus Harrison & Walker 1977 – cuculiform, coraciiform, close to Primapus?
†Pulchrapollia Dyke & Cooper 2000 – psittaciform ?
*†P. gracilis Dyke & Cooper 2000
*†P. olsoni
Neornithes incerta sedis USNM 496384 – parvicuculid? aegithalornithid?
†Palaeopsittacus georgei Harrison 1982b – caprimulgiform or quercypsittid?
†Amitabha urbsinterdictensis Gulas-Wroblewski & Wroblewski 2003 - galliform or gruiform ?
†Eociconia sangequanensis Hou 1989 – ciconiiform ?
†Protocypselomorphus manfredkelleri Mayr 2005 – caprimulgiform, apodiform or ancestral to both
†Pumiliornis tessellatus Mayr 1999
†Ludiortyx hoffmanni Brunet 1970 – rallid, quercymegapodid?
†Minggangia changgouensis Hou 1982 – rallid, threskiornithid?
†Petropluvialis simplex Harrison & Walker 1976 – may be same as Palaeopapia; anseriform?
†"Phasianus" alfhildae Shufeldt 1915 – gruiform, ciconiiform, phoenicopteriform?
†Telecrex grangeri Wetmore 1934 – meleagrid or gruiform
Neornithes incerta sedis AMNH FR 2941 – falconiform ? gruiform ?
†Zheroia kurochkini Nesov 1988 – gruiform? pelagornithid?
†"Falco" falconellus Shufeldt 1915 – falconiform ?
†Agnopterus Milne-Edwards 1868 – phoenicopteriform or anseriform
*†A. laurillardi Milne-Edwards 1868
*†A. sicki de Alvarenga 1990
*†A. turgaiensis Tugarinov 1940
†Plesiocathartes Gaillard 1908 - cathartid, leptosomid?
*†P. europaeus Gaillard 1908
*†P. gaillardi
*†P. wyomingensis Weidig 2006
*†P. major Weidig 2006
*†P. geiselensis Mayr 2002
*†P. kelleri Mayr 2002
†Botauroides parvus Shufeldt 1915 – coliiform?
†Aminornis excavatus Ameghino 1899 – gruiform ?
†Ciconiopsis antarctica Ameghino 1899 – ciconiiform ?
†Climacarthrus incompletus Ameghino 1899 – falconiform ? A nomen dubium
†Cruschedula revola Ameghino 1899 Aves incertae sedis; A nomen dubium
†Dolichopterus viator Milne-Edwards 1867 – charadriiform ? Not Dolicopterus as sometimes claimed
†Loncornis erectus Ameghino 1899 – gruiform ?
†Loxornis clivus Ameghino 1894– anatid?
†Manu antiquus Marples 1946 – pelagornithid? procellariiform ?
†Palaeocrex rex Wetmore 1927 – gruiform ?
†Palaeopapia eous > Harrison & Walker 1979 – anseriform?
†Paracygnopterus scotti Harrison & Walker 1979 – anseriform ?
†"Pararallus" hassenkampi Martini 1967
†Riacama caliginea Ameghino 1899 – gruiform?
†Smiliornis penetrans Ameghino 1899 – gruiform?
†Teracus littoralis Milne-Edwards 1871
†Teleornis impressus Ameghino 1899 – anatid?
†Pseudolarus guaraniticus Ameghino 1899 – gruiform?
Neornithes incerta sedis BMNH PAL 4989 – formerly "Ptenornis" and included in Headonornis; anseriform?
†"Anas" creccoides van Beneden 1871 – anseriform?
†"Charadrius" sheppardianus Cope 1881 – charadriiform
†Megagallinula harundinea Kuročkin 1968
†"Palaeorallus" alienus Kuročkin 1968 – galliform?
†"Vanellus" selysii van Beneden 1871 – charadriiform ?
†Anserpica kiliani Mourer-Chauviré, Berthet & Hugueney 2004 – gruiform or anseriform ?
†Gnotornis Wetmore 1942 – gruiform ?
*†G. aramiellus Wetmore 1942
*†G. roardeola
*†G. walkeri
†Guguschia nailiae Aslanova & Burčak-Abramovič 1968 – anseriform ? pelagornithid ?
†Tiliornis senex Ameghino 1899 – phoenicopteriform? A nomen dubium
Neornithes incerta sedis QM F40203 – gruiform ?
†Gaviella pusilla Shufeldt 1915 – gaviiform? plotopterid?
†"Anas" skalicensis Bayer 1882 - anseriform?
†Chenornis graculoides Portis 1884 - Anseriformes or Pelecaniformes ?
†"Propelargus" olseni Brodkorb 1963 – ciconiiform?
Neornithes incerta sedis MNHN SA 1259-1263 – passeriform?
†Anisolornis excavatus Ameghino 1891 – gruiform, galliform, tinamiform?
†"Ardea" perplexa Milne-Edwards 1869 – ardeid? strigiform?
†"Cygnus herrenthalsi" van Beneden 1871
†"Anas" risgoviensis Ammon 1918 – anseriform?
†"Ardea" aureliensis – ardeid?
†Eoneornis australis Ameghino 1895 nomen dubium – anatid? A nomen dubium
†Eutelornis patagonica Ameghino 1895 – anatid?
†Protibis cnemialis Ameghino 1891 – ciconiiform ?
†"Limnatornis" paludicola Milne-Edwards 1871 – coliid? phoeniculid?
†"Picus" gaudryi – piciform?
†"Ardea" lignitum Giebel 1860 – ardeid? strigid ?
†Bathoceleus hyphalus Brodkorb 1959 – picid?
†"Homalopus" Milne-Edwards 1870 non Chevrolat 1837– piciform? Preoccupied by a subgenus of Cryptocephalus leaf beetles described in 1835.
†"Liptornis hesternus" Ameghino 1894 – pelecaniform ? A nomen dubium
†Proceriavis martini Harrison & Walker 1979 – pelagornithid?
†Protopelicanus cuvieri Reichenbach 1852 – pelecaniform ? pelagornithid?
†Eurofluvioviridavis robustipes Mayr 2005a
†Archaeotrogonidae Wetmore 1926 – basal Cypselomorphae?
*†Archaeotrogonidae gen. et sp. indet.
*†Hassiavis laticauda Mayr 1998
*†Archaeotrogon Milne-Edwards 1892
†Cimolopterygidae Brodkorb 1963 – charadriiform?
*†Cimolopteryx Marsh 1892
**†C. rara Marsh 1890
**†C. maxima Brodkorb 1963
*†Lamarqueavis Agnolin 2010
**†L. australis Agnolin 2010
**†L. minima Agnolin 2010
**†L. petra Agnolin 2010
†Cladornithidae Wetmore 1930
†Eremopezidae Rothschild 1911 – pelecaniform? ratite?
*†Eremopezus eocaenus Andrews 1904 - includes Stromeria fajumensis Lambrecht 1929
†Gracilitarsidae Mayr 2001 – close to Sylphornithidae?
*Eutreptodactylus itaboraiensis Baird & Vickers-Rich 1997 - a nomen dubium
*Gracilitarsus mirabilis Mayr 1998
Halcyornithidae Harrison & Walker 1972 – psittaciform, coraciiform?
*†Halcyornis toliapicus Owen 1846
*†Cyrilavis Martin 2010 - psittaciform ?
**†C. olsoni Martin 2010
**†C. colburnorum Ksepka, Clarke & Grande 2011
†Juncitarsidae Peters 1987
*†Kashinia magnum Harrison & Walker 1979 – phoenicopteriform?
*†Juncitarsus Olson & Feduccia 1980 – phoenicopteriform?
Laornithidae Cracraft 1973 – charadriiform? gruiform? pelagornithid?
*Laornis edvardsianus Marsh 1870
†Lonchodytidae Brodkorb 1963
*†Lonchodytes estesi Brodkorb 1963 – gaviiform/pelecaniform? procellariiform?
†Palaeospizidae – passeriform? coraciiform? coliiform?
*†Palaeospiza bella Allen 1878
†Parvicuculidae Harrison 1982 – cypselomorph, cuculiform, coraciiform ?
*†Parvicuculus minor Harrison & Walker 1977
†Remiornithidae Martin 1992 – palaeognath?
*†Remiornis heberti Lemoine 1881
†Sylphornithidae Mourer-Chauviré 1988 – cuculiform? coraciiform? close to Gracilitarsidae?
*†Sylphornis bretouensis Mourer-Chauvire 1988
*†Oligosylphe mourerchauvireae Mayr & Smith 2002
†Tytthostonychidae Olson & Parris 1987 – procellariiform, pelecaniform?
*†Tytthostonyx glauconiticus Olson & Parris 1987
†Zygodactylidae Brodkorb 1971 – near passerine
*†Eozygodactylus americanus Weidig 2010
*†Zygodactylus Ballmann 1969
**†Z. ignotus Ballmann 1969
**†Z. luberonensis Mayr 2008
**†Z. grivensis Ballmann 1969
*†Primoscens minutus Harrison & Walker 1977
*†Primozygodactylus Mayr 1998
**†P. ballmanni Mayr 1998
**†P. danielsi Mayr 1998
**†P. major Mayr 1998
**†P. eunjooae Mayr & Zelenkov 2009
†"Graculavidae" Fürbringer 1888 – a paraphyletic form taxon, the "transitional shorebirds"
*†Limosavis Shufeldt 1915 – charadriiform?
**†L. velox Lambrecht 1933
**†L. augustus
*†Palaeotringa Marsh 1870 – charadriiform?
*†Telmatornis priscus Marsh 1870 – charadriiform? gruiform? podicipediform?
*†Zhylgaia aestiflua Nesov 1988 – presbyornithid?
*†Scaniornis lundgreni Dames 1890 – phoenicopteriform?
*†Dakotornis cooperi Erickson 1975
*†Placement unresolved
**"Graculavidae" gen. et sp. indet.
[Struthioniformes]

and related ratites.

Struthionidae Vigors 1825
*†Palaeotis weigelti Lambrecht 1928
*†Remiornis heberti Lemoine 1881
*†Orientornis linxiaensis Wang 2008
*Prehistoric species of extant genera
**Struthio Linnaeus 1758
[Casuariiformes]

, emus and related ratites.

Casuariidae Kaup 1847
*†Diogenornis fragilis de Alvarenga 1983 – possible casuariiform
*†Hypselornis sivalensis Lydekker
*†Emuarius Boles 1992
*Prehistoric species of extant genera
**Dromaius Vieillot 1816
**Casuarius Brisson 1760 non Reichenbach 1853 non Illiger 1826; Cela Moehring 1752 nom. rej.; Rhea Lacépède 1800 non Latham 1790; Chelarga Billberg 1828; Oxyporus Brookes 1828 non Donk 1933; Thrasys Billberg 1828; Cassowara Perry 1811; Hippalectryo Gloger 1842]
[Rheiformes]

and related ratites.

†Opisthodactylidae Ameghino 1895
*†Opisthodactylus Ameghino 1895 – rheid?
Rheidae
*†Heterorhea dabbenei Rovereto 1914
*†Hinasuri nehuensis Tambussi 1995
*Prehistoric species of extant genera
**Rhea Brisson 1760
†[Dinornithiformes]
†Megalapterygidae -
*†Megalapteryx didinus Haast 1886
†Dinornithidae Owen 1843
*†Dinornis Owen 1843
†Emeidae
*†Anomalopteryx didiformis Reichenbach 1853
*†Emeus crassus Reichenbach 1853
*†Euryapteryx Haast 1874
*†Pachyornis Lydekker 1891

Two unnamed Saint Bathans Fauna species.

[Apterygiformes]

Apterygidae Gray 1840 (Kiwis
*†Proapteryx micromeros Worthy et al. 2013
†[Lithornithiformes] Houde 1988
†Lithornithidae Houde 1988
*†Fissuravis weigelti Mayr 2007
*†Pseudocrypturus cercanaxius Houde 1988
*†Paracathartes howardae Harrison 1979
*†Lithornis Owen 1840
[Tinamiformes]
Tinamidae Gray 1840
*†Querandiornis romani Rusconi 1958
*†Roveretornis Brodkorb 1961
*Tinamidae gen. et sp. indet. MACN-SC Fleagle Collection - at least 2 species
*Prehistoric species of extant genera
**Eudromia
**Nothura Wagler 1827
**Crypturellus Brabourne & Chubb 1914
[Vegaviiformes]
†Neogaeornis wetzeli Lambrecht 1929
†Polarornis gregorii Chatterjee 2002
†Vegavis iaai Clarke et al. 2005
†Maaqwi cascadensis McLachlan, Kaiser, & Longrich, 2017
†Australornis lovei Mayr & Scofield, 2014
[Anseriformes]

The group that includes modern ducks and geese.

Basal and unresolved forms
*†Anatalavis Olson & Parris 1987 – anseranatid or basal.
**†A. rex Olson & Parris 1987
**†A. oxfordi Olson 1999
*†Romainvillia stehlini Lebedinský 1927 – anseranatide or anatide
*†Saintandrea chenoides Mayr & De Pietri 2013
*†Proherodius oweni Lydekker 1891 – presbyornithid?
*†Paranyroca magna Miller & Compton 1939 – anatide or own family?
*†Garganornis ballmanni Meijer 2014
Anhimidae Stejneger 1885
*†Bullockornis planei Rich 1979
*†Ilbandornis Rich 1979
**†I. woodburnei Rich 1979
**†I. lawsoni Rich 1979
*†Dromornis Owen 1872
**†D. australis Owen 1872
**†D. stirtoni Rich 1979
Anseranatidae Sclater 1880 sensu Stejneger 1885
*Anseranatidae gen. et sp. nov
†Presbyornithidae Wetmore 1926
*†Teviornis gobiensis Kuročkin, Dyke & Karhu 2002
*†Headonornis hantoniensis Harrison & Walker 1976 – only BMNH PAL 30325 belongs to this bird, may belong to Presbyornis.
*†Presbyornis Wetmore 1926
**†P. isoni Olson 1994
**†P. mongoliensis Kuročkin & Dyke 2010
**†P. recurvirostrus Ericson 2000
**†P. pervetus Wetmore 1926
*†Telmabates Howard 1955
**†T. antiquus Howard 1955
**†T. howardae Cracraft 1970
*†Wilaru Boles et al. 2013
**†W. tedfordi Boles et al. 2013
**†W. prideauxi Vanesa et al. 2016
Anatidae Leach 1820
*†Cygnavus
**†C. senckenbergi Lambrecht 1931
**†C. formosus Korochkin 1968
*†Cygnopterus Lambrecht 1931 – sometimes included in Cygnavus
**†C. neogradiensis Kessler & Hir 2009
**†C. affinis Lambrecht 1931
*†Mionetta Livezey & Martin 1988
**†M. robusta Mlíkovský 2002
**†M. arvernensis
**†M. blanchardi Livezey & Martin 1988
**†M. consobrina Livezey & Martin 1988
**†M. natator Livezey & Martin 1988
*†Australotadorna alecwilsoni Worthy 2009
*†Pinpanetta Worthy 2009
**†P. tedfordi Worthy 2009
**†P. vickersrichae Worthy 2009
**†P. fromensis Worthy 2009
*†Dunstanetta johnstoneorum Worthy et al. 2007
*†Manuherikia Worthy et al. 2007
**†M. lacustrina Worthy et al. 2007
**†M. minuta Worthy et al. 2007
**†M. douglasi Worthy et al. 2008
*†Matanas enrighti Worthy et al. 2007
*†Miotadorna sanctibathansi Worthy et al. 2007
*†Megalodytes morejohni Howard 1992
*†Sinanas diatomas Yeh 1980
*†Dendrochen Miller 1944
**†D. integra Cheneval 1987
**†D. oligocaena
**†D. robusta Miller 1944
*†Presbychen abavus Wetmore 1930
*†Afrocygnus chauvireae Louchart et al. 2005
*†Balcanas pliocaenica – may belong in Tadorna
*†Wasonaka yepomerae Howard 1966
*†Paracygnus plattensis Short 1969
*†Anabernicula Ross 1935
**†A. minuscula
**†A. oregonensis Howard 1964b
**†A. gracilenta Ross 1935
*†Eremochen russelli Brodkorb 1961
*†Tirarinetta kanunka Worthy 2008
*†Brantadorna Howard 1964
**†B. downsi Howard 1964
**†B. robusta Livezey 1997
*†Nannonetta invisitata Campbell 1979
*†Aldabranas cabri Harrison & Walker 1978
*†Ankonetta larriestrai Cenizo & Agnolín 2010
*†Cayaoa bruneti Tonni 1979
*†Chenoanas Zelenkov 2012
**†Chenoanas asiatica Zelenkov et al. 2018
**†Chenoanas deserta Zelenkov 2012
**†Chenoanas sansaniensis
*†Helonetta brodkorbi Emslie 1992
*†Mioquerquedula minutissima Zelenkov & Kuročkin 2012
*†Proanser major Umanskaya 1979
*†Shiriyanetta hasegawai Watanabe & Matsuoka 2015
*†Pleistoanser bravardi Agnolín 2006
*†Lavadytis pyrenae Stidham & Hilton 2015
*†Asiavis phosphatica Nesov 1986
*†Nogusunna conflictoides Zelenkov 2011
*†Sharganetta mongolica Zelenkov 2011
*†Protomelanitta Zelenkov 2011
**†P. gracilis Zelenkov 2011
**†P. shihuibas Zelenkov 2012
*†Bambolinetta lignitifila Mayr & Pavia 2014
*†Heteroanser vicinus Zelenkov 2012
*Placement unresolved
**†"Anas" albae Jánossy 1979 – formerly in Mergus
**†"Anas" amotape Campbell 1979
**†"Anas" eppelsheimensis Lambrecht 1933
**†"Anas" isarensis Lambrecht 1933
**†"Anas" luederitzensis
**†"Anas" sanctaehelenae Campbell 1979
**†"Anser" scaldii
**†"Aythya" chauvirae – 2 species
**†"Chenopis" nanus De Vis 1905 – at least 2 taxa, may be living species
**†"Cygnopterus" alphonsi Cheneval 1984
**†"Oxyura" doksana Mlíkovský 2002
**†"cf. Megalodytes"
**†Anatidae gen. et sp. indet. MNZ S42797
**†Anatidae gen. et spp. indet.
**†Anatidae gen. et spp. indet.
*Prehistoric species of extant genera
**Biziura Stephens 1824
***†B. exhumata DeVis 1889
***†B. delautouri Forbes 1892
**Dendrocygna
***†D. eversa Wetmore 1924
***†D. valdipinnis DeVis 1888
***†D. soporata Mlíkovský & Švec 1986
**Somateria
**Bucephala
***†B. cereti Boeuf & Mourer-Chauviré 1992
**Clangula
***†C. matraensis Kessler 2009
**Aix
***†A. praeclara Zelenkov & Kuročkin 2012
**Cygnus
***†C. mariae Bickart 1990
***†C. csakvarensis Mlíkovský 1992
***†C. liskunae
***†C. atavus Mlíkovský 1992
***†C. paloregonus Cope 1878
***†C. hibbardi Brodkorb 1958
***†C. lacustris
***†C. verae Boev 2000
**Chloephaga
***†C. robusta Tambussi 1998
**Histrionicus
***†Histrionicus shotwelli
**Mergus
***†M. connectens Jánossy 1972
***†M. minor Kessler 2009
***†M. miscellus Alvarez & Olson 1978
**Anas
***†A. elapsa DeVis 1888
***†A. gracilipes DeVis 1905
***†A. moldovica
***†A. strenuum
***†A. kisatibiensis
***†A. lambrechti
***†A. itchtucknee McCoy 1963
***†A. kurochkini Zelenkov & Panteleyev 2015
***†A. pullulans
***†A. schneideri Emslie 1985
***†A. bunkeri
***†A. greeni
***†A. ogallalae
***†A. cheuen Agnolín 2006
**Anser
***†A. arenosus Bickart 1990
***†A. arizonae Bickart 1990
***†A. azerbaidzhanicus Serebrovsky 1940
***†A. devjatkini Kuročkin 1971
***†A. djuktaiensis Zelenkov & Kuročkin 2014
***†A. eldaricus Burchak-Abramovich & Gadzyev 1978
***†A. oeningensis Milne-Edwards 1867b
***†A. pratensis
***†A. tchikoicus Kuročkin 1985
***†A. thompsoni Martin & Mengel 1980
***†A. thraceiensis Burchak-Abramovich & Nikolov 1984
***†A. pressa
***†A. udabnensis Burchak-Abramovich 1957
**Aythya
***†A. chauvirae Cheneval 1987
***†A. denesi Zelenkov 2016
***†A. effodiata
***†A. magna Kuročkin 1985
***†A. molesta Zelenkov 1985
***†A. reclusa
***†A. robusta
***†A. spatiosa Kuročkin 1976
**Branta
***†B. dickey Miller 1924
***†B. esmeralda Burt 1929
***†B. howardae Miller 1930
***†B. hypsibata
***†B. propinqua Schufeldt 1892
***†B. thessaliensis Boev & Koufos 2006
**Oxyura
***†O. bessomi Howard 1963
***†O. hulberti Emslie 1992
***†O. zapatanima Alvarez 1977
**Chendytes
***†O. milleri Howard 1955
**Lophodytes
**Neochen
***†N. barbadiana Brodkorb 1965
***†N. debilis
***†N. pugil
**Tadorna
***†T. minor Kessler & Hir 2012
***†T. petrina Kuročkin 1985
[Galliformes]

The group that includes domestic chickens and their relatives.

Placement unresolved
*†Archaealectrornis sibleyi Crowe & Short 1992 - phasianid?
*†Archaeophasianus Lambrecht 1933 - tetraonid or phasianid
**†A. mioceanus
**†A. roberti
*†Austinornis lentus Clarke 2004 - tentatively placed here;
*†Chambiortyx cristata Mourer-Chauviré et al. 2013
*†"Cyrtonyx" tedfordi
*†Linquornis gigantis Yeh 1980
*†Namaortyx sperrgebietensis Mourer-Chauviré, Pickford & 2011
*†Palaealectoris incertus Wetmore 1930 - tetraonid?
*†Palaeoalectoris songlinensis Hou 1987
*†Palaeonossax senectus Wetmore 1956 - cracid?
*†Palaeortyx Milne-Edwards 1869 - phasianid or odontophorid
**†P. media Milne-Edwards 1871
**†P. volans Göhlich & Pavia 2008
**†P. gallica Milne-Edwards 1869 non Lydekker 1891
**†P. brevipes Milne-Edwards 1869 emend. Paris 1912
**†P. prisca
**†P. phasianoides Milne-Edwards 1869
**†P. joleaudi Ennouchi 1930
*†Procrax brevipes Tordoff & Macdonald 1957 - cracid? gallinuloidid?
*†Shandongornis Yeh 1997
**†S. shanwanensis Yeh 1997
**†S. yinansis Hou 2003
*†Sobniogallus albinojamrozi Tomek et al. 2014
*†Taoperdix Milne-Edwards 1869 - gallinuloidid?
**†T. pesseiti
**†T. miocaena Ballmann 1969
*Galliformes gen. et sp. indet. MCZ 342506 - formerly in Gallinuloides; phasianid?
†Gallinuloididae Lucas 1900
*†Gallinuloides wyomingensis Eastman 1900
*†Paraortygoides Mayr 2000
**†P. messelensis Mayr 2000
**†P. radagasti Dyke & Gulas 2002
†Paraortygidae Mourer-Chauviré 1992
*†Pirortyx major Brodkorb 1964
*†Scopelortyx klinghardtensis Mourer-Chauviré, Pickford & Senut 2015
*†Paraortyx Gaillard 1908 sensu Brodkorb 1964
**†P. lorteti Gaillard 1908
**†P. brancoi Gaillard 1908
†Quercymegapodiidae Mourer-Chauviré 1992
*†Taubacrex granivora de Alvarenga 1988
*†Ameripodius de Alvarenga 1995
**†A. silvasantosi Alvarenga 1995
**†A. alexis Mourer-Chauviré 2000
*†Quercymegapodius Mourer-Chauviré 1992
**†Q. depereti
**†Q. brodkorbi Mourer-Chauviré 1992
Megapodidae – megapodes
*†Ngawupodius minya Boles & Ivison 1999
*Prehistoric species of extant genera
**†Leipoa gallinacea – formerly Chosornis, Palaeopelargus, Progura
Cracidae – guans and curassows
*†Boreortalis laesslei Brodkorb 1954 – may be same as Ortalis
*Extant genera present in the fossil record
**Ortalis
***†O. affinis Feduccia & Wilson 1967
***†O. phengites Wetmore 1923
***†O. pollicaris Miller 1944
***†O. tantala Wetmore 1933
Odontophoridae – New World quails
*†Nanortyx inexpectatus Weigel 1963
*†Neortyx peninsularis Holman 1961
*†Miortyx Miller 1944
**†M. teres Miller 1944
**†M. aldeni Howard 1966
*Placement unresolved
**Odontophoridae gen. et sp. indet. KUVP 9393
*Prehistoric species of extant genera
**†Cyrtonyx cooki Gutierrez et al. 1981
**†Callipepla shotwelli
**†Colinus hibbardi Wetmore 1944
**†Colinus suilium Brodkorb 1959
**†Colinus sp.
**†Dendrortyx? sp.
Phasianidae – pheasants, quails, partridges, grouse and turkeys
*†Bantamyx georgicus Kuročkin 1982
*†Diangallus mious Hou 1985
*†Lophogallus naranbulakensis Zelenkov & Kuročkin 2010
*†Megalocoturnix cordoni Sánchez Marco 2009
*†Miophasianus Brodkorb 1952
**†M. maxima Brodkorb 1964
**†M. altus Villalta & Crusafont 1950
**†M. medius Lambrecht 1933
*†Proagriocharis kimballensis Martin & Tate 1970
*†Palaeocryptonyx Depéret 1892
**†P. donnezani Depéret 1892
**†P. grivensis Ennouchi 1930
**†P. novaki Sánchez Marco 2009
**†P. edwardsi Ballmann 1969a
**†P. minor
**†P. subfrancolinus
*†Pliogallus coturnoides Běljajeva 1948
*†Plioperdix Kretzoi 1955
**†P. crassipes
**†P. kormosi
**†P. africana Mourer-Chauviré & Geraads 2010
**†P. hungarica Zelenkov & Panteleyev 2014
*†Rustaviornis georgicus Burchak-Abramovich & Meladze 1972
*†Rhegminornis calobates Wetmore 1943
*†Schaubortyx keltica Brodkorb 1964
*†Shandongornis Yeh 1997
**†S. shanwanensis Yeh 1997
**†S. yinansis Hou 2003
*†Shanxiornis fenyinis Wang et al. 2006
*†Tologuica Zelenkov & Kuročkin 2009
**†T. aurorae Zelenkov & Kuročkin 2009
**†T. karhui Zelenkov & Kuročkin 2009
*Placement unresolved
**Tetraoninae gen. et sp. indet.
**Meleagridae gen. et sp. indet.
**"Tympanchus" stirtoni Miller 1944
**"Tympanuchus" lulli Shuefeldt 1915
*Extant genera present in the fossil record
**Coturnix
**Bambusicola
***†B. dalianensis
***†B. nini Sánchez Marco 2009
***†B. praebonasia Bocheński 1991
**Phasianus
***†P. lufengia Hou 1985
***†P. yanshansis Huang & Hou 1984
**Crossoptilon
***†C. jiai Hou 1982
**Lophura
***†L. wayrei Harrison & Walker 1982
**Gallus
***†G. aesculapii Jánossy 1976
***†G. beremendensis Jánossy 1977
***†G. georgicus
***†G. imereticus
***†G. karabachensis Baryšnikov & Potapova 1995
***†G. kudarensis Burčak-Abramovič & Potapova
***†G. meschtscheriensis
***†G. tamanensis
**Lagopus
***†L. atavus Jánossy 1974
***†L. balcanicus Boev 1995
***†L. lagopus noaillensis Mourer-Chauviré 1975
***†L. muta correzensis Mourer-Chauviré 1975
**Meleagris
***†M. celer Marsh 1872
***†M. tridens Wetmore 1931
***†M. progenes
***†M. leopoldi
***†M. anza
***†M. crassipes Rea 1980
**Pavo
***†P. bravardi
**Tetrao
***†T. conjugens Jánossy 1974
***†T. macropus Jánossy 1976
***†T. partium
***†T. praeurogallus Jánossy 1969
***†T. rhodopensis Boev 1998
**Lyrurus
***†Lyrurus tetrix longipes
**Francolinus
***†F. capeki Jánossy 1974
***†F. villanyiensis Jánossy 1974
***†F. wenzensis Jánossy 1974
**Perdix
***†P. inferna
***†P. margaritae Kuročkin 1985
***†P. palaeoperdix Mourer-Chauviré 1975
**Syrmaticus
***†S. kozlovae Kuročkin 1985
**Bonasa
***†B. dalianensis
***†B. nini Sánchez Marco 2009
***†B. praebonasia Bocheński 1991
**Dendragapus
***†D. gilli
***†D. lucasi Jehl 1969
**Alectoris
***†A. baryosefi Tchernov 1980
***†A. bavarica Ballmann 1969
***†A. peii'' Hou 1982
[Charadriiformes]

s, auks, shorebirds

Basal and unresolved taxa
*Charadriiformes gen. et sp. indet. – burhinid? basal?
*"Morsoravis" – a nomen nudum?
*Jiliniornis – charadriid?
*Boutersema – glareolid?
*Turnipax – turnicid?
*Elorius
*"Larus desnoyersii – larid? stercorarid?
*"Larus pristinus – larid?
*Charadriiformes gen. et spp. indet. – several species, 1 probably larid
*Charadriiformes gen. et sp. indet.
*"Totanus" teruelensis – scolopacid? larid?
*"Actitis" balcanica – scolopacid? charadriid?
Scolopacidae – waders and snipes
*Paractitis
*Mirolia
*Placement unresolved
**Scolopacidae gen. et sp. indet.
**Scolopacidae gen. et sp. indet.
*Extant genera present in the fossil record
**Limosa
**Tringa – includes Totanus
**Gallinago
**Scolopax
**Phalaropus
**Actitis
**Numenius – includes Palnumenius
Jacanidae – jacanas
*Nupharanassa
*Janipes
*Prehistoric species of extant genera
**Jacana farrandi
Laridae – gulls
*Laridae gen. et sp. indet.
*Laricola – larid? Formerly "Larus" elegans and "L." totanoides
*Gaviota Miller & Sibley 1941 non Fischer 1983
*Extant genera present in the fossil record
**Larus
Alcidae – auks
*Hydrotherikornis
*Pseudocepphus
*Petralca
*Miocepphus
*Alcodes
*Praemancalla
*Mancalla
*Extant genera present in the fossil record
**Cepphus
**Cerorhinca
**Uria
**Aethia
**Alca
**Synthliboramphus
**Fratercula
**Pinguinus
**Brachyramphus
**Ptychoramphus
Stercorariidae – skuas and jaegers
*Prehistoric species of extant genera
**Stercorarius sp.
**Stercorarius shufeldti
**Prehistoric subspecies of extant species
***Stercorarius pomarinus philippi
Glareolidae – pratincoles
*Paractiornis
*Mioglareola - formerly "Larus" dolnicensis
*Prehistoric species of extant genera
**Glareola neogena
Burhinidae – thick-knees
*Prehistoric species of extant genera
**Burhinus lucorum
**Burhinus aquilonaris
**Burhinus sp.
**Burhinus sp.
**Prehistoric subspecies of extant species
***Burhinus bistriatus nanus
Charadriidae – plovers
*Limicolavis
*Viator – may be synonym of Vanellus
*Extant genera present in the fossil record
**Vanellus – includes Belanopteryx
*Additional prehistoric species of extant genera
**Oreopholus orcesi
Recurvirostridae – avocets
*Extant genera present in the fossil record
**Himantopus
*Additional prehistoric species of extant genera
**Recurvirostra sanctaeneboulae
†[Gastornithiformes] (extinct)

The diatrymas, a group of huge flightless Paleogene birds of unclear affinities. Traditionally placed within the Gruiformes, they are usually considered a distinct order nowadays and appear closer to the Anseriformes.

Gastornithidae
*†Ornitholithes biroi Dughi & Sirugue 1969
*†Ornithoformipes controversus
*†Rivavipes giantess
*†Gasthornithidae gen. et sp. indet. – possibly Gastornis
*†Gasthornithidae gen. et sp. indet. YPM PU 13258 – possibly juvenile Gastornis giganteus
*†"Diatryma" cotei Galliard 1936
*†Gastornis Hébert 1855
**†G. parisiensis Hébert 1855
**†G. giganteus
**†G. sarasini Mlíkovský 2002
**†G. russeli Martin 1992
**†G. xichuanensis Buffetaut 2013
[Gruiformes]

The group that includes modern rails and cranes. Probably paraphyletic.

Placement unresolved
*†Rupelrallus Fischer 1997 – rallid? parvigruid?
**†R. belgicus Mayr 2012
**†R. saxoniensis Fischer 1997
*†"Gruiformes" gen. et sp. indet. MNZ S42623 – Aptornithidae?
†Songziidae Hou 1990 possibly a nomen nudum
*†Songzia Hou 1990 possibly a nomen nudum
**†S. heidangkouensis Hou 1990
**†S. acutunguis Wang et al. 2012
†Eogruidae Wetmore 1934

*†Sonogrus gregalis Kuročkin 1981
*†Eogrus Wetmore 1932
**†E. aeola Wetmore 1932
**†E. crudus Kuročkin 1981
**†E. turanicus
**†E. wetmorei Brodkorb 1967
*†Ergilornithinae
*†Proergilornis minor Kozlova 1960
*†Ergilornis Kozlova 1960
**†E. rapidus
**†E. minor
*†Amphipelargus Lydekker 1891
**†A. brodkorbi
**†A. majori Lydekker 1891
**†A. dzabghanensis Kuročkin 1985
**†A. cracrafti Kuročkin 1985
**†A. ukrainus Kuročkin 1985
**†A. orientalis Kuročkin 1985
†Geranoididae Wetmore 1933
*†Eogeranoides campivagus Cracraft 1969
*†Geranoides jepseni Wetmore 1933
*†Geranodornis aenigma Cracraft 1969
*†Galligeranoides boriensis Bourdon, Mourer-Chauviré & Laurent 2014
*†Palaeophasianus Shufeldt 1913
**†P. meleagroides Shufeldt 1913
**†P. incompletus Cracraft 1969
*†Paragrus Lambrecht 1933
**†P. prentici Lambrecht 1933
**†P. shufeldti Cracraft 1969
†Parvigruidae Mayr 2005
*†Parvigrus pohli Mayr 2005
Aramidae Bonaparte 1854
*†Badistornis aramus Wetmore 1940 – aramid?
*Extant genera present in the fossil record
**Aramus paludigrus Rasmussen 1997 - Aramid
Gruidae – cranes
*†Camusia quintanai Seguí 2002
*†Aramornis Wetmore 1926 – gruid? aramid?
**†A. longurio Wetmore 1926
**†A. robalearica
**†A. crataegensis
*†Palaeogrus Portis 1885
**†P. princeps Portis 1884
**†P. hordwelliensis Lambrecht 1933
**†P. mainburgensis Göhlich 2003
*†Geranopsis hastingsiae Lydekker 1871 Hordwell Late Eocene– Early Oligocene of England)– gruiform or anseriform ?
*†Eobalearica tugarinovi Gureev 1949
*†Pliogrus
*†Probalearica mongolica Kurochkin 1985 gruid? A nomen dubium?
*Gruidae gen. et sp. indet. – formerly Grus conferta Miller & Sibley 1942
*Extant genera present in the fossil record
**Balearica
***†B. crataegensis
***†B. excelsa Mlíkovský 2002
***†B. exigua Feduccia & Voorhies 1992
***†B. rummeli Mourer-Chauviré 2001
**Grus
***†G. bogatshevi
***†G. haydeni Marsh 1870
***†G. latipes
***†G. moldavica Mlíkovský 2002
***†G. nannodes Wetmore & Martin 1930
***†G. pagei Campbell 1995
***†G. pentelici Gaudry 1862
***†G. primigenia Milne-Edwards 1869
***†G. melitensis Lydekker 1890
Rallidae – rails
*†Aletornis Marsh 1872
**†A. marshi Lambrecht 1933
**†A. bellus Marsh 1872
**†A. gracilis Marsh 1872
**†A. nobilis Marsh 1872
**†A. pernix Marsh 1872
*†Australlus Worthy & Boles 2011
**†A. disneyi Worthy & Boles 2011
**†A. gagensis Worthy & Boles 2011
*†Baselrallus intermedius De Pietri & Mayr 2014
*†Belgirallus Mayr & Smith 2001
**†B. minutus Mayr & Smith 2001
**†B. oligocaenus Mayr & Smith 2001
*†Creccoides osbornii Shufeldt 1892
*†Eocrex Wetmore 1931
**†E. primus Wetmore 1931
**†E. tagusevae Panteleyev 2001
*†Euryonotus Mercerat 1897 – rallid?
**†E. uspallatensis Mercerat 1897
**†E. brachypterus Mercerat 1897
**†E. argentinus Mercerat 1897
*†Fulicaletornis venustus Lambrecht 1933
*†Ibidopsis hordwelliensis Lydekker 1891
*†Latipons Harrison & Walker 1979
**†L. gardneri Harrison & Walker 1979
**†L. robinsoni Harrison & Walker 1979
*†Miofulica dejardini Lambrecht 1933
*†Miorallus major Lambrecht 1933
*†Palaeoaramides Lambrecht 1933
**†P. lungi Olson 1977
**†P. minutus Cracraft 1973
**†P. tugarinovi Kuročkin 1980
**†P. christyi Lambrecht 1933
**†P. minor
**†P. beaumontii Cracraft 1973
*†Palaeorallus Wetmore 1931
**†P. troxelli Wetmore 1931
**†P. brodkorbi Cracraft 1973
*†Paraortygometra porzanoides Lambrecht 1933
*†Parvirallus Harrison & Walker 1979
**†P. gracilis Harrison & Walker 1979
**†P. bassetti Harrison 1984
**†P. gassoni Harrison 1984
**†P. medius Harrison 1984
*†Pastushkinia zazhigini Zelenkov 2013
*†Pleistorallus flemingi Worthy 1997
*†Quercyrallus
**†Q. ludianus Brodkorb 1963
**†Q. arenarius Lambrecht 1933
**†Q. quercy Cracraft 1973
*†Rallicrex
**†R. litkensis Kessler & Hir 2012
**†R. polgardiensis Jánossy 1991
**†R. kolozsvarensis Lambrecht 1933
*†Rhenanorallus rhenanus Mayr 2010
*†Wanshuina lii Hou 1994
*†Youngornis
**†Y. gracilis Yeh 1981
**†Y. qiluensis Yeh & Bo 1989
*Placement unresolved
**Rallidae gen. et sp. indet.
**Rallidae gen. et spp. indet. – several species
**Rallidae gen. et sp. indet.
**Rallidae gen. et sp. indet.
**Rallidae gen. et sp. indet. UMMP V55013/-14; UMMP V55012/V45750/V45746
**Rallidae gen. et sp. indet. UMMP V29080
**Rallidae gen. et sp. indet.
*Extant genera present in the fossil record
**Rallus
***†R. auffenbergi
***†R. cyanocavi Steadman & Wright 2013
***†R. ibycus Olson & Wingate 2000
***†R. lacustris Brodkorb
***†R. minutus Alcover et al. 2015 non Pallas 1776 non Gmelin 1789 non Forster 1844
***†R. natator
***†R. phillipsi Wetmore
***†R. prenticei Wetmore 1944
***†R. recessus Olson & Wingate 2001
***†R. richmondi Olson 1977
**Porzana
***†P. botunensis Boev 2015
***†P. estramosi Jánossy 1979
***†P. kretzoii Kessler 2009
***†P. matraensis Kessler 2009
***†P. piercei Olson & Wingate 2000
***†P. risillus Kuročkin 1980
**Gallinula
***†G. balcanica Boev 1999
***†G. brodkorbi McCoy 1963
***†G. gigantea Černov 1980
***†G. kansarum Brodkorb 1967
**Fulica
***†F. infelix Brodkorb 1961
***†F. podagrica Brodkorb 1965
***†F. stekelesi Tchernov 1968
**Laterallus
***†L. guti Brodkorb 1952
***†L. insignis Feduccia 1968
***†L. sp.
**Coturnicops
***†C. avita Feduccia 1968
**Porphyrio
***†P. parvus'' Harrison & Walker 1982
[Eurypygiformes]
†Aptornithidae Bonaparte 1856
*†Aptornis proasciarostratus Worthy, Tennyson & Scofield 2011
Eurypygidae Selby 1840
*†Eoeurypyga olsoni Weidig 2003 nomen nudum
†Messelornithidae Hesse 1988 – Messel-birds
*†Pellornis mikkelseni Bertelli, Chiappe & Mayr 2011
*†Itardiornis hessae Mourer-Chauviré 1995
*†Messelornis Hesse 1988
**†M. russelli Mourer-Chauviré 1995
**†M. nearctica Hesse 1992
**†M. cristata Hesse 1988
[Cariamiformes]
†Itaboravis elaphrocnemoides Mayr, Alvarenga & Clarke 2011
†Walbeckornis creber Mayr 2007
†Qianshanornithidae Mayr et al. 2013
*†Qianshanornis rapax Mayr et al. 2013
†Salmilidae Mayr 2002
*†Salmila robusta Mayr 2000
†Ameghinornithidae Mourer-Chauviré 1981
*†Strigogyps Gaillard 1908
**†S. dubius Gaillard 1908
**†S. robustus
**†S. sapea
†Bathornithidae Wetmore 1933
*†Eutreptornis uintae Cracraft 1971
*†Neocathartes grallator Wetmore 1950
*†Paracrax Brodkorb 1964
**†P. antiqua Cracraft 1971
**†P. wetmorei Cracraft 1968
**†P. gigantea Cracraft 1968
*†Bathornithidae gen. nov. – formerly Bathornis celeripes Wetmore 1933 and B. cursor Wetmore 1933
*†Bathornis Wetmore 1927
**†B. minor Cracraft 1971
**†B. veredus Wetmore 1927
**†B. fax Cracraft 1968
**†B. geographicus Wetmore 1942
**†B. fricki Cracraft 1968

†Idiornithidae Brodkorb 1965
*†Dynamopterus Milne-Edwards 1892
**†I. cursor Oberholser 1899
**†I. gallicus Oberholser 1899
**†I. gracilis Mourer-Chauviré 1983
**†I. minor Oberholser 1899
**†D. anthracinus Mourer-Chauviré 2013
**†D. tuberculata Mourer-Chauviré 2013
**†D. gaillardi Mourer-Chauviré 2013
**†D. itardiensis Mourer-Chauviré 2013
*†Talantatos Reichenbach 1852
**†T. bordkorbi Mlíkovský 2002
**†T. crex
**†T. phasianus
**†T. fossilis Reichenbach 1852
**†T. proudlocki Mlíkovský 2002
*†Gypsornis Milne-Edwards 1869
**†G. cuvieri Milne-Edwards 1869
**†G. venablesi Mlíkovský 2002
*†Propelargus Lydekker 1891
**†P. edwardsi
**†P. insolitus Mlíkovský 2002
**†P. cayluxensis Lydekker 1891
*†Occitaniavis elatus Mourer-Chauvire 1983
*†Ibidopodia Milne-Edwards 1868 – threskiornithid?
**†I. chavrochensis De Pietri & Mayr 2014
**†I. minuta De Pietri & Mayr 2014
**†I. palustris Milne-Edwards 1868
†Phorusrhacidae Ameghino 1899 – Terror birds
*†Lavocatavis africana Mourer-Chauviré et al. 2011
*†Patagorhacos terrificus Agnolin & Chafrat 2015
*†Eleutherornis Schaub 1940
**†E. helveticus Schaub 1940
**†E. cotei Gaillard 1936
*†Paleopsilopterus itaboraiensis de Alvarenga 1985
*†Procariama simplex Rovereto 1914
*†Psilopterus Moreno & Mercerat 1891
**†P. bachmanni
**†P. lemoinei
**†P. affinis
**†P. colzecus Tonni & Tambussi 1988
*†Llallawavis scagliai Degrange et al. 2015
*†Mesembriornis Moreno 1889
**†M. milneedwardsi Moreno 1889
**†M. incertus
*†Patagornis marshi Moreno & Mercerat 1891
*†Andrewsornis abbotti Paterson 1941
*†Andalgalornis steulleti Patterson & Kraglievich 1960
*†Phorusrhacos longissimus Ameghino 1887
*†Devincenzia pozzi Kraglievich 1932
*†Titanis walleri Brodkorb 1963
*†Kelenken guillermoi Bertelli, Chiappe & Tambussi 2007
*†Physornis fortis Ameghino 1895
*†Paraphysornis brasiliensis de Alvarenga 1993
*†Brontornis burmeisteri Moreno & Mercerat 1891
Cariamidae Bonaparte 1850 – seriemas
*†Riacama caliginea Ameghino 1899
*†Pseudolarus guaraniticus Ameghino 1899
*†Smiliornis penetrans Ameghino 1899
*†Noriegavis santacrucensis Agnolin 2009
*Prehistoric species of extant genera
**Chunga
***†C. incerta Tonni 1974
[Otidiformes]
Otididae – bustards
*†Gryzaja odessana Zubareva 1939
*†Ioriotis gabunii Burchak-Abramovich & Vekua 1981
*†Miootis compactus Umanskaya 1979
*†Pleotis liui Hou 1982
*Prehistoric species of extant genera
**Chlamydotis Brodkorb 1967
***†C. affinis
***†C. mesetaria Sánchez Marco 1990
**Tetrax
***†T. paratetrax
**Otis
***†O. bessarabicus Kessler & Gal 1996
***†O. hellenica Boev, Lazaridis & Tsoukala 2014
***†O. khosatzkii Bocheński & Kuročkin 1987
[Phoenicopteriformes]
Placement unresolved
*†Phoeniconotius eyrensis Miller 1963
†Palaelodidae Stejneger 1885 – Swimming-flamingos
*†Adelalopus hoogbutseliensis Mayr & Smith 2002 -
*†Palaelodus Milne-Edwards 1863
**†P. ambiguus Milne-Edwards 1863
**†P. aotearoa Worthy et al. 2010
**†P. germanicus
**†P. kurochkini Zelenkov 2013
**†P. pledgei Baird & Vickers-Rich 1998
**†P. wilsoni Baird & Vickers-Rich 1998
*†Megapaloelodus Miller 1944
**†M. conectens
**†M. peiranoi Agnolin 2009
**†M. goliath Cheneval 1983c
**†M. opsigonus Brodkorb 1961
Phoenicopteridae – flamingos
*†Harrisonavis croizeti Kashin 1978
*†Leakeyornis aethiopicus Vickers-Rich & Walker 1983
*†Elornis Milne-Edwards 1868
**†E. anglicus Aymard 1856 nomen nudum
**†E. grandis Milne-Edwards 1868
**†E. littoralis Milne-Edwards 1868
*Placement unresolved
**Phoenicopteridae gen. et sp. indet.
*Extant genera present in the fossil record
**Phoenicopterus
***†P. floridanus Brodkorb 1953
***†P. stocki
***†P. siamensis Cheneval et al. 1991
***†P. gracilis Miller 1963
[Podicipediformes]
Podicipedidae – grebes
*†Miobaptus Švec 1982
**†M. huzhiricus Zelenkov 2015
**†M. walteri Švec 1982
*†Miodytes serbicus Dimitreijevich, Gál & Kessler 2002
*†Pliolymbus baryosteus Murray 1967
*†Thiornis sociata Navás 1922
*Placement unresolved
**Podicipedidae gen. et sp. indet. – formerly included in Podiceps parvus
**Podicipedidae gen. et sp. indet. UMMP 49592, 52261, 51848, 52276, KUVP 4484
**Podicipedidae gen. et sp. indet.
*Extant genera present in the fossil record
**Aechmophorus
***†A. elasson Murray 1967
**Podilymbus
***†P. mujusculus Murray 1967
***†P. wetmorei Storer 1976
***†P. podiceps magnus
**Podiceps
***†P. arndti Chandler 1990
***†P. caspicus
***†P. csarnotatus Kessler 2009
***†P. discors Murray 1967
***†P. dixi Brodkorp 1963
***†P. miocenicus Kessler 1984
***†P. oligocaenus
***†P. parvus
***†P. solidus Kuročkin 1985
***†P. subparvus
[Phaethontiformes]
†Prophaethontidae Harrison & Walker 1976
* †Lithoptila abdounensis Bourdon, Bouya & Iarochène 2005 – including Abdounornis
* †Prophaethon shrubsolei Andrews 1899
Phaethontidae Brandt 1831 – tropicbirds
*?†Proplegadis fisheri Harrison & Walker 1971 – Threskiornithidae?
* †Phaethusavis pelagicus Bourdon, Amaghzaz & Bouya 2008
* †Heliadornis Olson 1985
** †H. ashbyi Olson 1985
** †H. minor Kessler 2009
** †H. paratethydicus Mlíkovský 1997
[Ciconiiformes]

The diverse group that includes storks, herons and New World vultures. Paraphyletic as listed here.

†Ciconiidae – storks
* †Ciconiopsis antarctica Ameghino 1899
* †Palaeoephippiorhynchus dietrichi Lambrecht 1930
* †Grallavis edwardsi Cheneval 1984 – may be same as Prociconia
* †Pelargosteon tothi Kretzoi 1962
* †Sanshuiornis zhangi Wang et al. 2012
*Placement unresolved
**Ciconiidae gen. et sp. indet. – formerly Aquilavus bilinicus Brodkorb 1964; Cygnus bilinicus Laube 1909
**Ciconiidae gen. et sp. indet.
**cf. Leptoptilos gen. et sp. indet. – formerly Leptoptilos siwalicensis Harrison 1974
**Ciconiidae gen. et sp. indet. – Ciconia or Mycteria?
*Extant genera present in the fossil record
**Ciconia – includes Xenorhynchus
*** †C. lydekkeri Ameghino 1891
*** †C. gaudryi Lambrecht 1933
*** †C. minor Harrison 1980
*** †C. sarmatica Grigorescu & Kessler 1977
*** †C. kahli Haarhoff 1988
*** †C. louisebolesae Boles 2005
*** †C. lucida Kuročkin 1982
*** †C. maltha Miller 1910
*** †C. nana Rich & van Tets 1982
*** †C. stehlini Janossy 1992
**Mycteria
*** †M. milleri
*** †M. wetmorei Howard 1935
**Ephippiorhynchus
*** †E. tchoufour Louchart et al. 2008
*** †E. pakistanensis Harrison & Walker 1982
**Leptoptilos – includes Cryptociconia
*** †L. falconeri Lydekker 1884
*** †L. pliocenicus Zubareva 1948
*** †L. inidcus
*** †L. lüi Zhang et al. 2012
*** †L. patagonicus Noriega & Cladera 2008
*** †L. robustus Meijer & Awe Due 2010
*** †L. richae Harrison 1974
*** †L. titan Wetmore 1940
**Jabiru
*** †J. codorensis Walsh & Sánchez 2008
[Pelecaniformes]

The group that includes modern pelicans and cormorants. As presented here paraphyletic; the tropicbird lineage is not part of this group and relationships with Procellariiformes and Sphenisciformes require more research. Also, as the pelicans are at least as close to the Ciconiiformes as to cormorants, the latter group is being recognized as Phalacrocoraciiformes by some recent authors and the core Pelecaniformes are occasionally merged into the Ciconiiformes.

Basal and unresolved forms
* †Piscator tenuirostris Harrison & Walker 1976 – basal phalacrococacoid?
* †"Prophalacrocorax ronzoni Harrison 1975
* † "Pelecaniformes" gen. et sp. indet. basal phalacrococacoid ?
Fregatidae – frigatebirds
* †Limnofregata Olson 1977
** †L. azygosternon Olson 1977
** †L. hutchisoni Stidham 2014
** †L. hasegawai Olson & Matsuoka 2005
Sulidae – gannets and boobies
* †Bimbisula melanodactylus Benson & Erickson 2013
* †Eostega lebedinskyi Lambrecht 1929
* †Empheresula arvenensis Harrison 1975
* †Masillastega rectirostris Mayr 2002
* †Microsula pygmaea Wetmore 1938
* †Miosula media Miller 1925
* †Palaeosula stocktoni Howard 1958
* †Rhamphastosula Stucchi & Urbina 2004
** †R. ramirezi Stucchi & Urbina 2004
** †R. aguirrei Stucchi, Varas-Malca & Urbina-Schmitt 2015
* †Sarmatosula dobrogensis Grigorescu & Kessler 1977
* Placement unresolved
** Sulidae gen. et sp. indet.
** Sulidae gen. et sp. indet.
* Extant genera present in the fossil record
** Morus
*** †M. avitus
*** †M. lompocanus Domning 1978
*** †M. loxostylus Brodkorb 1963
*** †M. olsoni Grigorescu & Kessler 1988
*** †M. peninsularis Brodkorb 1955
*** †M. peruvianus Stucchi 2003
*** †M. recentior Howard 1949
*** †M. reyanus Howard 1936
*** †M. vagabundus Wetmore 1930
*** †M. willetti
** Sula
*** †S. brandi Stucchi, Varas-Malca & Urbina-Schmitt 2015
*** †S. figueroae Stucchi, Varas-Malca & Urbina-Schmitt 2015
*** †S. clarki Chandler 1990
*** †S. humeralis Miller & Bowman 1958
*** †S. magna Stucchi 2003
*** †S. pohli Howard 1958
*** †S. sulita Stucchi 2003
*** †S. universitatis Brodkorb 1963

Phalacrocoracidae – cormorants and shags

* †Borvocarbo Mourer-Chauviré, Berthet & Hugueney 2004 – phalacrocoracid? basal phalacrococacoid?
** †B. guilloti Mourer-Chauviré, Berthet & Hugueney 2004
** †B. tardatus Göhlich & Mourer-Chauviré 2010
* †Limicorallus Kuročkin 1968
** †L. saiensis Kuročkin 1968
** †L. carbunculus Mayr 2009
* †Nambashag Worthy 2011
** †N. billerooensis Worthy 2011
** †N. microglaucus Worthy 2011
* †Nectornis Cheneval 1984
** †N. africanus Mayr 2014
** †N. miocaenus
** †N. anatolicus Mlíkovský 1998
* †Valenticarbo praetermissus Harrison 1979 – a nomen dubium
* †Oligocorax Lambrecht 1933
** †O. littoralis Milne-Edwards 1868
** †O. stoeffelensis Mayr 2014
* Placement unresolved
** †Phalacrocoracidae gen. et sp. indet.
* Extant genera present in the fossil record
** Phalacrocorax – may be several genera. Includes Australocorax, Miocorax
*** †P. gregorii De Vis 1905
*** †P. gregorii
*** †P. chapalensis Alvarez 1977
*** †P. destefani
*** †P. femoralis
*** †P. filyawi Emslie 1995b
*** †P. goletensis Howard 1965
*** †P. ibericus Villalta 1963
*** †P. idahensis
*** †P. intermedius
*** †P. kennelli
*** †P. kuehneanus Schlüter 1991
*** †P. longipes
*** †P. leptopus Brodkorb 1961
*** †P. macer Brodkorb
*** †P. macropus
*** †P. marinavis
*** †P. mongoliensis Kuročkin 1971
*** †P. owrei Brodkorb & Mourer-Chauviré 1984
*** †P. reliquus Kuročkin 1976
*** †P. rogersi
*** †P. serdicensis Burčak-Abramovič & Nikolov 1984
*** †P. tanzaniae Harrison & Walker 1979
*** †P. wetmorei Brodkorb 1955
†Plotopteridae – diving-"boobies"
* †Copepteryx Olson & Hasegawa 1996
** †C. hexeris Olson & Hasegawa 1996
** †C. titan Olson & Hasegawa 1996
* †Hokkaidornis abashiriensis Sakurai, Kimura & Katoh 2008
* †Phocavis maritimus Goedert 1988
* †Plotopterum joaquinensis Howard 1969
* †Tonsala Olson 1980
** †T. hildegardae Olson 1980
** †T. buchanani Dyke, Xia & Habib 2011
†Protoplotidae
* †Protoplotus beauforti Lambrecht 1931
Anhingidae – darters
* †Meganhinga chilensis Alvarenga 1995
* †Macranhinga Noriega 1992
** †M. paranensis Noriega 1992
** †M. ranzii de Alvarenga & Guilherme 2003
* †"Paranavis" – a nomen nudum
* †Giganhinga kiyuensis Inderknecht & Noriega 2002
* Extant genera present in the fossil record
** Anhinga
*** †A. beckeri Emslie 1998
*** †A. hadarensis Brodkorb & Mourer-Chauviré 1982
*** †A. laticeps
*** †A. malagurala Mackness 1995
*** †A. pannonica Lambrecht 1916
*** †A. subvolans Becker 1986
*** †A. walterbolesi Worthy 2012
*** †A. fraileyi Campbell 1996
*** †A. grandis Martin & Mengel 1975
*** †A. minuta de Alvarenga & Guilherme 2003
Pelecanidae – pelicans
* †Protopelicanus cuvieri Reichenbach 1852
* †Miopelecanus Cheneval 1984
** †M. gracilis Cheneval 1984
** †M. intermedius Cheneval 1984
* Extant genera present in the fossil record
** Pelecanus
*** †P. aethiopicus Harrison & Walker 1976
*** †P. cautleyi Davies 1880
*** †P. fraasi Lydekker 1891
*** †P. halieus Wetmore 1933
*** †P. odessanus Widhalm 1886
*** †P. schreiberi Olson 1999
*** †P. sivalensis Davies 1880
*** †P. tirarensis Miller 1966

†Pelagornithidae Fürbringer 1888 – pseudotooth birds
* †Pseudodontornis Lambrecht 1930
** †P. longidentata Harrison & Walker 1976
** †P. tenuirostris Harrison 1985
** †P. tschulensis
** †P. longirostris Lambrecht 1930
** †P. stirtoni Howard & Warter 1969
* †Dasornis Owen 1870
** †D. abdoun Bourdon 2010
** †D. emuinus Owen 1870
** †D. gigas
* †Macrodontopteryx
* †Odontopteryx toliapica Owen 1873
* †Gigantornis eaglesomei Andrews 1916
* †Cyphornis magnus Cope 1894
* †Osteodontornis orri Howard 1957
* †Pelagornis
** †P. miocaenus Lartet 1857
** †P. mauretanicus Mourer-Chauviré & Geraads 2008
** †P. chilensis Mayr & Rubilar-Rogers 2010
** †P. sandersi Ksepka 2014
* †Caspiodontornis kobystanicus Aslanova & Burčak-Abramovič 1982
* †Tympanoneisiotes wetmorei Hopson 1964
* †Lutetodontopteryx tethyensis Mayr & Zvonok 2012
* †Aequornis traversei Bourdon 2006 nomen nudum
* Placement unresolved
** ?Pelagornithidae gen. et sp. indet.
†Xenerodiopidae Rasmussen 1987
* †Xenerodiopus mycter Rasmussen, Olson & Simons 1987
Ardeidae – herons
* †Ardeagrandis arborea Kurockin & Ganya 1972
* †Calcardea junnei Gingerich 1987
* †Matuku otagoense Scofield, Worthy & Tennyson 2010
* †Nyctisoma robusta Elżanowski & Zelenkov 2015
* †Palaeophoyx columbiana McCoy 1963
* †Pikaihao bartlei Worthy et al. 2013
* †Proardea amissa Lambrecht 1933
* †Proardeola walkeri Harrison 1979) – possibly same as Proardea
* †Zeltornis ginsburgi Balouet 1981
*Placement unresolved
** †Anas basaltica Bayer 1882
*Extant genera present in the fossil record
** Nyctanassa
*** †N. kobdoena Kuročkin 1976
** Ardea
*** †A. aurelianensis Milne-Edwards 1871
*** †A. polkensis Brodkorb 1955
*** †A. howardae Brodkorb 1980
*** †A. paloccidentalis Shufeldt 1892
*** †A. sytchevskayae Zelenkov 2011
** Egretta
*** †E. polgardiensis Kessler 2009
*** †E. subfluvia Becker 1985
** Butorides
*** †B. validipes Campbell 1976
** Botaurus
*** †B. hibbardi Moseley & Feduccia 1975
** Syrigma
*** †S. sanctimartini Campbell 1979
Scopidae – hammerkop
*Prehistoric species of extant genera
** †Scopus xenopus Olson 1984
Threskiornithidae – ibises
*Threskiornithidae gen. et sp. indet. NMMP-KU 1301 – Threskiornithinae?
* †Gerandibis paganus De Pietri 2013
* †Rhynchaeites messelensis Wittich 1898 – may include Mopsitta
* †Vadaravis brownae Smith, Grande & Clarke 2013
*Extant genera present in the fossil record
**Geronticus
*** †G. perplexus Cheneval 2000
*** †G. apelex Olson 1985
*** †G. olsoni Mourer-Chauviré & Geraads 2010
**Platalea
*** †P. chione
*** †P. tiangangensis Hou 1987
**Plegadis
*** †P. pharangites Olson 1981
**Theristicus
*** †T. wetmorei Campbell 1979
**Eudocimus
*** †E. leiseyi Emslie 1995a
*** †E. peruvianus Campbell 1979
*** †Eudocimus sp.
Balaenicipitidae – shoebills
*Goliathia andrewsi Lambrecht 1930
*Paludiavis richae Harrison & Walker 1982
[Procellariiformes]

The group that includes modern albatrosses, petrels and storm-petrels.

Placement unresolved
*†Eopuffinus kazachstanensis Nesov 1986
*†Makahala mirae Mayr 2015
*†Proaestrelata Imber 1985
†Marinavidae Harrison & Walker 1977
*†Marinavis longirostris Harrison & Walker 1977
†Tytthostonychidae Olson & Parris 1987
*†Tytthostonyx glauconiticus Olson & Parris 1987
†Diomedeoididae Fischer 1985
*†Rupelornis van Beneden 1871
**†R. harmati Mayr & Smith 2012
**†R. definitus van Beneden 1871
**†R. babaheydariensis Mayr & Smith 2012
Oceanitidae
*Extant genera present in the fossil record
**Oceanites
***†O. zaloscarthmus Olson 1985
Diomedeidae – albatrosses
*†Murunkus subitus Panteleyev & Nesov 1993
*†Plotornis Milne-Edwards 1874
**†P. arvernensis Cheneval 1984a
**†P. deltortrii Milne-Edwards 1874
**†P. graculoides Mayr & Pavia 2014
*†Tydea septentrionalis Mayr & Smith 2012
*†Diomedeidae gen. et sp. indet.
*Extant genera present in the fossil record
**Diomedea
***†D. anglica Lydekker 1891a
***†D. howardae Chandler 1990
***†D. milleri Howard 1966
***†D. rumana Grigorescu & Kessler 1988
***†D. tanakai Davis 2003
***†D. thyridata Wilkinson 1969
**Phoebastria
***†P. rexsularum Olson & Rasmussen 2001
**Thalassarche
***†T. thyridata
Hydrobatidae – storm-petrels
*†Primodroma bournei Harrison & Walker 1977
*Prehistoric species of extant genera
**†"Oceanodroma" hubbsi Miller 1951
**†Pelagodroma sp. 1
**Pelagodroma sp. 2
Procellariidae – petrels
*†Argyrodyptes microtarsus Ameghino 1905
*†Pterodromoides minoricensis Sequí et al. 2001
*Extant genera present in the fossil record
**Calonectris
***†C. krantzi Olson & Rasmussen 2001
***†C. kurodai Olson 2009
***†C. wingatei Olson 2008
**Fulmarus
***†F. hammeri Howard 1968
***†F. miocaenus Howard 1984
**Pachyptila
***†P. salax Olson 1985
**Pelecanoides
***†P. cymatotrypetes Olson 1985
***†P. miokuaka Worthy et al. 2007
***†P. urinatrix Gmelin 1789
**"Pterodroma"
***†P. kurodai Harrison & Walker 1978
***†P. rupinarum Olson 1975
**Puffinus
***†P. aquitanicus Brodkorb 1963 Procellaria aquitanica Milne-Edwards 1874
***†P. antiquus Brodkorb 1963c Procellaria antiqua Milne-Edwards 1874
***†P. barnesi Howard 1978
***†P. calhouni Howard 1968
***†P. conradi Marsh 1870
***†P. diatomicus Miller 1925
***†P. felthami Howard 1949
***†P. gilmorei Chandler 1990
***†P. inceptor Wetmore 1930
***†P. kanakoffi Howard 1949
***†P. micraulax Brodkorb 1963
***†P. mitchelli Miller 1961
***†P. nestori Alcover 1989
***†P. priscus Miller 1961
***†P. raemdonckii Brodkorb 1962
***†P. tedfordi Howard 1971
[Gaviiformes]
†Gaviella pusilla Wetmore 1940
Gaviidae – loons
*†Colymboides Milne-Edwards 1867 – paraphyletic?
**†C. harundinea
**†C. metzleri Mayr 2004
**†C. anglicus Lydekker 1891a
**†C. belgicus Mayr & Smith 2002
**†C. minutus Milne-Edwards 1867
* Placement unresolved
**"Gavia" portisi – rentatively placed here. A nomen dubium?
*Extant genera present in the fossil record
**Gavia
***†G. egeriana Švec 1982
***†G. brodkorbi Howard 1978
***†G. paradoxa Umanskaja 1981
***†G. schultzi Mlíkovsky 1998
***†G. howardae Brodkorb 1953c
***†G. moldavica Kessler 1984
***†G. concinna Wetmore 1940
***†G. fortis Olson & Rasmussen 2001
[Sphenisciformes]
Unresolved and basal forms
*†Waimanu Jones, Ando & Fordyce 2006
**†W. manneringi Jones, Ando & Fordyce 2006
**†W. tuatahi Jones, Ando & Fordyce 2006
*†Perudyptes devriesi Clarke et al. 2007 – basal?
*Sphenisciformes gen. et sp. indet. CADIC P 21
Spheniscidae – penguins
*†Kaiika maxwelli Fordyce & Tomas 2011
*†Palaeoapterodytes ictus Ameghino 1905
*†Tereingaornis moisleyi Scarlett 1983
*†Crossvallia unienwillia Tambussi et al. 2005
*†Anthropornis Wiman 1905
**†A. nordenskjoeldii Wiman 1905
**†A. grandis
*†Archaeospheniscus Marples 1952
**†A. lowei Marples 1952
**†A. lopdelli Marples 1952
*†Delphinornis Wiman 1905
**†D. gracilis Myrcha et al. 2002
**†D. larseni Wiman 1905
**†D. arctowskii Myrcha et al. 2002
**†D. wimani Ksepka & Clarke 2010
*†Palaeeudyptes Huxley 1859
**†P. klekowskii Myrcha, Tatur & del Valle 1990
**†P. gunnari
*†Icadyptes salasi Clarke et al. 2007
*†Inkayacu paracasensis Clarke et al. 2010
*†Pachydyptes ponderosus Oliver 1930
*†Marambiornis exilis Myrcha et al. 2002
*†Mesetaornis polaris Myrcha et al. 2002
*†Tonniornis Tambussi et al. 2006
**†T. mesetaensis Tambussi et al. 2006
**†T. minimum Tambussi et al. 2006
*†Wimanornis seymourensis Simpson 1971
*†Arthrodytes andrewsi Ameghino 1901
*†Duntroonornis parvus Marples 1953
*†Kairuku Ksepka et al. 2012
**†K. grebneffi Ksepka et al. 2012
**†K. waitaki Ksepka et al. 2012
*†Korora oliveri Marples 1952
*†Platydyptes Marples 1952
**†P. marplesi Simpson 1971
**†P. novaezealandiae Marples 1952
**†P. amiesi Marples 1952
*†Eretiscus tonnii Olson 1986
*†Palaeospheniscus Moreno & Mercerat 1891
**†P. huxleyorum Simpson 1973
**†P. wimani
**†P. gracilis Ameghino 1899
**†P. patagonicus Moreno & Mercerat 1891
**†P. bergi Moreno & Mercerat 1891
**†P. biloculata Acosta Hospitaleche 2007
*†Paraptenodytes Ameghino 1891
**†P. brodkorbi Simpson 1972
**†P. robustus
**†P. antarcticus Ameghino 1891
*†Anthropodyptes gilli Simpson 1959
*†Madrynornis mirandus Hospitaleche et al. 2007
*†Pseudaptenodytes Simpson 1970
**†P. minor Simpson 1970
**†P. macraei Simpson 1970
*†Dege hendeyi Simpson 1979
*†Marplesornis novaezealandiae Simpson 1972
*†Nucleornis insolitus Simpson 1979
*†Inguza predemersus Simpson 1975
*Placement unresolved
**Spheniscidae gen. et sp. indet
*Extant genera present in the fossil record
**Pygoscelis
***†P. calderensis Walsh & Suárez 2006
***†P. tyreei Simpson 1972
***†P. grandis Walsh & Suárez 2006
**Spheniscus
***†S. muizoni Göhlich 2007
***†S. chilensis Emslie & Correa 2003
***†S. megaramphus Stucchi, Urbina & Giraldo 2003
***†S. urbinai Stucchi 2002
**Aptenodytes
***†A. ridgeni Simpson 1972
**Eudyptes
***†E. calauina Hoffmeister et al. 2014
[Pterocliformes]
Pteroclidae – sandgrouse
*†Gerandia calcaria Lambrecht 1933
*†Leptoganga sepultus Mourer-Chauviré 1993
*†Archaeoganga Mourer-Chauviré 1992
**†A. validus
**†A. pinguis Mourer-Chauviré 1992
**†A. larvatus
*Extant genera present in the fossil record
**Syrrhaptes
***†S. kashini Kuročkin 1985
[Columbiformes]
Columbidae – doves and pigeons
*Arenicolumba – doubtfully distinct from Patagioenas
*Rupephaps
*Placement unresolved
**Columbidae gen. et sp. indet.
*Extant genera present in the fossil record
**Columba
**Patagioenas
[Psittaciformes]

Unresolved and basal fossil parrots:

Psittacopes
Serudaptus – pseudasturid or psittacid?
Pseudasturidae
Messelasturidae Mayr 2005
*Tynskya eocaena Mayr 2000
*Messelastur gratulator Peters 1994
*Pseudasturidae FU 125 gen. et sp. indet.
*Pseudasturides – formerly Pseudastur
Vastanavidae
*Vastanavis
Quercypsittidae
*Quercypsitta
Cacatuidae
*Extant genera present in the fossil record
**Cacatua
Psittacidae – parrots, parakeets and lories
*Archaeopsittacus
*Xenopsitta
*Psittacidae gen. et spp. indet. – several species
*Bavaripsitta
*Psittacidae gen. et sp. indet. – erroneously placed in Pararallus dispar, includes "Psittacus lartetianus
*Extant and recently extinct genera present in the fossil record
**Conuropsis
**Nandayus
**Cyanoliseus
**Aratinga
**Rhynchopsitta
Strigopidae – New Zealand parrots, kakapo
*Nelepsittacus
*Heracles
[Opisthocomiformes]

Hoatzins

Opisthocomidae Swainson 1837
*†Foro panarium Olson 1992 - cuculiform?
*†Hoatzi panarium Thomas 1996
*†Onychopteryx simpsoni Cracraft 1971 – falconid? A nomen dubium
*†Protoazin parisiensis Mayr & De Pietri 2014
*†Namibiavis senutae Mourer-Chauviré 2003
*†Hoazinavis lacustris Alvarenga, Mayr & Mourer-Chauviré 2011
*†Hoazinoides magdalenae Miller 1953
[Musophagiformes]
Musophagidae – turacos
*Placement unresolved
**Musophagidae gen. et sp. indet.
**Musophagidae gen. et sp. indet.
*†Veflintornis Kashin 1976
**†V. meini
**†V. africanus
[Cuculiformes]

Cuckoos, turacos and allies.

Placement unresolved
*Cuculiformes gen. et sp. indet.
Cuculidae – cuckoos
*Eocuculus
*Dynamopterus
*Neococcyx
*Cursoricoccyx
*Placement unresolved
**Cuculidae gen. et sp. indet.
[Accipitriformes]
†Teratornithidae Miller 1909– teratorns
*†Argentavis magnificens Campbell & Tonni 1980
*†Aiolornis incredibilis Campbell, Scott & Springer 1999
*†Oscaravis olsoni Suarez & Olson 2009
*†Cathartornis gracilis Miller 1910
*†Taubatornis campbelli Olson & Alvarenga 2002
*†Teratornis Miller 1909
**†T. woodburnensis Campbell & Stenger 2002
**†T. merriami Miller 1909
Cathartidae – New World vultures
*†Brasilogyps faustoi de Alvarenga 1985
*†Kuntur cardenasi Stucchi et al. 2015
*†Pleistovultur nevesi de Alvarenga et al. 2008
*†Tapinopus ellioti Milne-Edwards 1891
*†Diatropornis ellioti
*†Phasmagyps patritus Wetmore 1927
*†Hadrogyps aigialeus Emslie 1988
*†Pliogyps Becker 1986
**†P. charon Becker 1986
**†P. fisheri Tordoff 1959
*†Perugyps diazi Stucchi & Emslie 2005
*†Dryornis pampeanus – may belong to modern genus Vultur
*†Aizenogyps toomeyae Emslie 1998
*†Breagyps clarki Miller 1938
*†Geronogyps reliquus Campbell 1979
*†Wingegyps cartellei de Alvarenga & Olson 2004
*†Parasarcoramphus milneedwardsi Mourer-Chauviré 2002
*Placement unresolved
**Cathartidae gen. et sp. indet.
**Cathartidae gen. et sp. indet.
**Cathartidae gen. et sp. indet.
**Cathartidae gen. et sp. indet.
*Extant genera present in the fossil record
**Sarcoramphus
***†S. kernense
***†S. fischeri Campbell 1979
**Gymnogyps
***†G. howardae Campbell 1979
***†G. kofordi Emslie 1988
**Vultur – distinctiveness disputed
***†V. fossilis Moreno & Mercerat 1891
†Masillaraptoridae Mayr 2009
*†Masillaraptor parvunguis Mayr 2006
†Horusornithidae Mourer-Chauviré 1991
*†Horusornis vianeyliaudae Mourer-Chauviré 1991
Pandionidae – ospreys
*Extant genera present in the fossil record
**Pandion
***†P. lovensis Becker 1985
***†P. homalopteron Warter 1976
Sagittariidae – secretarybirds
*†Amanuensis pickfordi Mourer-Chauviré 2003
*†Pelargopappus Stejneger 1885 – formerly Amphiserpentarius/Amynoptilon/Pelargopsis
**†P. schlosseri
**†P. magnus
Accipitridae' – hawks, eagles and Old World vultures
*Milvoides
*Aquilavus
*Palaeocircus
*Palaeastur
*Pengana
*Promilio
*Proictinia
*Neophrontops – formerly in Neophron
*Mioaegypius
*Apatosagittarius
*Gansugyps
*Palaeoborus
*Qiluornis
*Thegornis
*Garganoaetus
*Amplibuteo – may belong to extant genus Harpyhaliaetus
*Neogyps
*Palaeohierax – includes "Aquila" gervaisii
*Placement unresolved
**Accipitridae gen. et sp. indet. AMNH FR 7434
**Accipitridae gen. et sp. indet.
**Accipitridae gen. et sp. indet.
**Accipitridae gen. et sp. indet. MPEF-PV-2523
**Accipitridae gen. et sp. indet. – Parabuteo?
**Accipitridae gen. et sp. indet. – Buteo?
**Accipitridae gen. et sp. indet.
**"Aquila" danana – formerly also Geranoaetus or Buteo
*Extant genera present in the fossil record
**Haliaeetus
**Buteo
**Aquila
**Buteogallus – might include Harpyhaliaetus
**"Hieraaetus" – doubtfully distinct from Aquila
**Milvus
**Gyps
**Aegypius
*Additional prehistoric species of extant genera
**Spizaetus grinnelli – formerly Geranoaetus or Buteo
**Spizaetus pliogryps – formerly Aquila
[Falconiformes]
Falconidae – falcons
*†Lagopterus minutus Moreno & Mercerat 1891
*†Parvulivenator watteli Harrison 1982
*†Stintonornis mitchelli Harrison 1984
*†Badiostes patagonicus Ameghino 1895
*†Pediohierax ramenta Becker 1987 -
*†Thegornis musculosus Ameghino 1895
**†Petrosushkinia pliocaena Zelenkov 2015
**†Falconidae gen. et sp. indet.
*Extant genera present in the fossil record
**Milvago
***†M. alexandri Olson 1976
***†M. brodkorbi Campbell 1979
**Falco
***†F. antiquus Mourer-Chauviré 1975
***†F. bakalovi Boev 1999
***†F. bulgaricus Boev 2011
***†F. chowi Hou 1982
***†F. hezhengensis Li et al. 2014
***†F. medius Umanskaya 1981
***†F. oregonus Brodkorb 1964
***†F. pisanus Portis 1887
***†F. umanskajae Sobolev 2003
[Steatornithiformes]
Paraprefica Mayr 1999
*†P. kelleri Mayr 1999
*†P. major Mayr 1999
Steatornithidae – oilbirds
*†Prefica nivea Olson 1987
*Prehistoric species of extant genera
**Steatornis sp.
Nyctibiidae – potoos
*†Euronyctibius kurochkini Mourer-Chauviré 1989
[Podargiformes]
†Fluvioviridavidae Mayr 2005
*†Fluvioviridavis platyrhamphus Mayr & Daniels 2001
Podargidae – frogmouths
*†Masillapodargus longipes Mayr 1999
*†Quercypodargus olsoni Mourer-Chauvire 1989
[Caprimulgiformes]
Caprimulgidae – nightjars
*†Ventivorus ragei Mourer-Chauviré 1988
*Prehistoric species of extant genera
**Caprimulgus
***†C. piurensis Campbell 1979
[Aegotheliformes]

Owlet-nightjars

Aegothelidae
*†Quipollornis koniberi Rich & McEvey 1977
*Extant genera present in the fossil record
**Aegotheles
***†A. savesi Layard & Layard 1881
[Apodiformes]

Swifts and hummingbirds.

†Wyomingcypselus pohli Weidig 2003 nomen nudum
†Procypseloides mourerchauvireae Harrison & Walker 1977
†Laputavis robusta Dyke 2001b
†Palescyvus escampensis Karhu 1988
†Scaniacypselus Harrison 1984
*†S. wardi Harrison 1984
*†S. szarskii Meyr & Peters 1999
†Aegialornithidae Lydekker 1891
*†Primapus lacki Harrison & Walker 1975 ?apodid
*†Aegialornis Lydekker 1891
**†A. leenhardti
**†A. germanicus Mlíkovský 2002
**†A. gallicus Lydekker 1891
**†A. wetmorei Collins 1976
**†A. broweri Collins 1976
†Eocypselidae Harrison 1984 – apodiform ? caprimulgiform? basal Cypselomorphae?
*†Eocypselus Harrison 1984
**†E. vincenti Harrison 1984
**†E. rowei Ksepka et al. 2013
†Cypselavidae Mourer-Chauviré 2006
*†Parargornis messelensis Mayr 2003 jungornithid, trochilid, basal as Argornis?
*†Argornis caucasicus Karhu 1999 ?Trochilidae
*†Cypselavus gallicus Gaillard 1908 aegialornithid or hemiprocnid
†Jungornithidae Karhu 1988
*†Jungornis Karhu 1988
**†J. tesselatus Karhu 1988
**†J. geraldmayri Mourer-Chauviré & Sigé 2006
Trochilidae hummingbirds
*†Eurotrochilus Mayr 2004
**†E. inexpectatus Mayr 2004
**†E. noniewiczi Bochenski & Bochenski 2008
*Placement unresolved
**Trochilidae sp. et gen. indet.
**Trochilidae sp. et gen. indet.
Apodidae – swifts
*Extant genera present in the fossil record
**Collocalia
**Apus
**Chaetura
**Tachornis
[Coliiformes]

Mousebirds and relatives

Unresolved and basal forms
* †Botauroides parvus Shufeldt 1915
* †Eobucco brodkorbi Feduccia & Martin 1976 - sandcoleid?
* †Eocolius walkeri Dyke & Waterhouse 2001 - sandcoleid or coliid
*†Limnatornis Milne-Edwards 1871 - coliid?
* †Coliiformes gen. et sp. indet.
*†Uintornis Marsh 1872 - sandcoleid?
Family †Chascacocoliidae Zelenkov & Dyke 2008
* Genus †Chascacocolius Houde & Olson 1992 - basal? sandcoleid?
Family †Selmeidae Zelenkov & Dyke 2008
*†Selmes absurdipes Peters 1999 - coliid?
Family †Sandcoleidae Houde & Olson 1992 sensu Mayr & Mourer-Chauviré 2004
*†Sandcoleus copiosus Houde & Olson 1992
*†Anneavis anneae Houde & Olson 1992
*†Eoglaucidium pallas Fischer 1987
Family Coliidae Swainson 1837 sensu Mayr & Mourer-Chauviré 2004
*†Primocolius Mourer-Chauviré 1988
*†Oligocolius Mayr 2000
*†Masillacolius brevidactylus Mayr & Peters 1998
*Extant genera present in the fossil record
** Colius
[Strigiformes]

Owls and barn owls

Unresolved and basal forms
*Berruornis – basal? Sophornithidae?
*Strigiformes gen. et sp. indet.
*Palaeoglaux – own family Palaeoglaucidae or Strigidae?
*Palaeobyas – Tytonidae? Sophiornithidae?
*Palaeotyto – Tytonidae?
*Strigiformes gen. et spp. indet.
Ogygoptyngidae
*Ogygoptynx
Protostrigidae
*Eostrix
*Minerva – formerly Protostrix, includes "Aquila" ferox, "Aquila" lydekkeri, and "Bubo" leptosteus
* Oligostrix
Sophiornithidae
*Sophiornis
Strigidae – typical owls
*Mioglaux – includes "Bubo" poirreiri
*Intulula – includes "Strix/Ninox" brevis
*Alasio – includes "Strix" collongensis
*Oraristrix
*Placement unresolved
**"Otus" wintershofensis – may be close to extant genus Ninox
**"Strix" edwardsi
**"Asio" pygmaeus
**Strigidae gen. et sp. indet. UMMP V31030 – Strix/Bubo?
*Extant genera present in the fossil record
**Strix
**Bubo
**Asio
**Athene
**Glaucidium
**Surnia
**Pulsatrix
Tytonidae – barn owls
*Nocturnavis
*Selenornis
*Necrobyas
*Prosybris
*Placement unresolved
**Tytonidae gen. et sp. indet. TMT 164
*Extant genera present in the fossil record
**Tyto
[Coraciiformes]

Rollers and allies. Probably paraphyletic.

Basal and unresolved forms
*†Quasisyndactylus longibrachis Mayr 1998 – alcediniform, basal?
*†Paracoracias occidentalis Clarke et al. 2009
*†Cryptornis antiquus Gervais 1848–52 – bucerotid? geranopterid?
*†Protornis glarniensis von Meyer 1856 – basal to motmotids and meropids?
*† Coraciiformes gen. et spp. indet. PQ 1216, QU 15640 – 2 species
†Geranopteridae Mayr & Mourer-Chauviré 2000
*†Geranopterus Milne-Edvards 1892
**†G. alatus
**†G. bohemicus Mourer-Chauvire 1999
**†G. milneendwardsi Mayr & Mourer-Chauvire 2000
†Eocoraciidae Mayr & Mourer-Chauviré 2000
*†Eocoracias brachyptera Mayr & Mourer-Chauviré 2000
†Primobucconidae Feduccia & Martin 1976 sensu Mayr, Mourer-Chauviré & Weidig 2004
*†Primobucco Brodkorb 1970
**†P. kestneri Feduccia 1973
**†P. frugilegus Mayr, Mourer-Chauviré & Weidig 2004
**†P. mcgrewi Brodkorb 1970
**†P. perneri Mayr, Mourer-Chauviré & Weidig 2004
Todidae – todies
*†Palaeotodus Olson 1976
**†P. emryi Olson 1976
**†P. escampsiensis Mourer-Chauviré 1985
**†P. itardiensis Mourer-Chauviré 1985
Motmotidae – motmots
*Placement unresolved
**Momotidae gen. et sp. indet.
Coraciidae
*†Miocoracias chenevali Mourer-Chauviré, Peyrouse & Hugueney 2013
*Extant genera present in the fossil record
**Eurystomus
***†E. beremedensis Kessler 2010
[Bucerotiformes]
†Messelirrisoridae
*†Messelirrisor Mayr 1998
**†M. halcyorostris Mayr 1998
**†M. parvus Mayr 1998
**†M. grandis Mayr 2000
Phoeniculidae
*†Phirriculus pinicola Mlíkovský & Göhlich 2000
Bucerotidae – hornbills
*†Euroceros bulgaricus Boev & Kovachev 2007
*Extant genera present in the fossil record
**Bucorvus
***†B. brailloni Brunet 1971
[Trogoniformes]
Trogonidae – trogons
*†Foshanornis songi Zhao et al. 2015
*†Septentrogon madseni Kristoffersen 2002
*†Paratrogon gallicus Lambrecht 1933
*†Primotrogon Mayr 1999
**†P. wintersteini Mayr 1999
**†P. pumilio Mayr 2005
*Placement unresolved
**Trogonidae gen. et sp. indet. 1
**Trogonidae gen. et sp. indet. 2
[Piciformes]
Placement unresolved
*Piciformes gen. et sp. indet. IRScNB Av 65
*Rupelramphastoides – ramphastid?
*Piciformes gen. et sp. indet. SMF Av 429
*Capitonides – ramphastid? "capitonid" ? own family Capitonididae?
*Pici gen. et sp. indet. – "capitonid"
Miopiconidae
*Miopico
Lybiidae – African barbets
*Lybiidae gen. et sp. indet. – extant genus Pogoniulus?
Galbulidae
*Extant genera present in the fossil record
**Galbula hylochoreutes
Picavidae
*Picavus
Picidae – woodpeckers
*Palaeopicus
*Palaeonerpes
*Pliopicus
*Placement unresolved
**Picidae gen. et sp. indet.
**Picidae gen. et sp. indet.
**cf. Colaptes DMNH 1262
*Extant genera present in the fossil record
**Campephilus
**Colaptes
**Dendrocopos
**Additional prehistoric subspecies of extant species
***Melanerpes superciliaris ssp.
***Melanerpes superciliaris ssp.
[Passeriformes]
Placement unresolved
*Passeriformes gen. et spp. indet. – several species, oscine?
*Resoviaornis
*Wieslochia
*Passeriformes gen. et spp. indet. – several suboscine and oscine species
*Certhiops – basal Certhioidea
*Passeriformes gen. et sp. indet. – suboscine?
*Passeriformes gen. et spp. indet. – several species, oscine?
*Passeriformes gen. et spp. indet. – several species, basal?
*Passeriformes gen. et spp. indet. – several species, oscine?
*Passeriformes gen. et spp. indet. – Sylvioidea
*"Palaeostruthus" eurius
Eurylaimidae – broadbills
*Placement unresolved
**Eurylaimidae gen. et sp. indet.
Palaeoscinidae
*Palaeoscinis
Furnariidae – ovenbirds
*Pseudoseisuropsis
**Pseudoseisuropsis nehuen
**Pseudoseisuropsis cuelloi
*Prehistoric species of extant genera
**Pseudoseisura cursor
**Cinclodes major
Menuridae – lyretails
*Extant genera present in the fossil record
**Menura
Meliphagidae – honeyeaters
*Placement unresolved
**Meliphagidae gen. et spp. indet. – at least 7 spp., some may be from extant genera
Orthonychidae – logrunners
*Extant genera present in the fossil record
**Orthonyx
Oriolidae – Old World orioles
*Longmornis
Artamidae – woodswallows, butcherbirds, currawongs and Australian magpie
*Placement unresolved
**Artamidae gen. et sp. indet. – cracticine
Corvidae – crows, ravens, jays and magpies
*Miocorvus
*Miopica
*Miocitta
*Protocitta
*Henocitta
*Extant genera present in the fossil record
**Corvus
**Pica
**Pyrrhocorax
*Placement unresolved
**Corvidae gen. et sp. indet.
**Corvidae gen. et sp. indet. – probably belongs in extant genus
Laniidae – shrikes
*Extant genera present in the fossil record
**Lanius
Regulidae – kinglets
*Extant genera present in the fossil record
**Regulus
Hirundinidae – swallows and martins
*Placement unresolved
**Hirundinidae gen. et spp. indet. – 2 species
Megaluridae – grass-warblers and allies
*Extant genera present in the fossil record
**?Locustella
Acrocephalidae – marsh- and tree-warblers
*Extant genera present in the fossil record
**?Acrocephalus
Muscicapidae – Old World flycatchers and chats
*Extant genera present in the fossil record
**Luscinia
Turdidae – thrushes
*Extant genera present in the fossil record
**?Turdus
Alaudidae – larks
*Eremarida
Motacillidae – wagtails
*Extant genera present in the fossil record
**Motacilla
Fringillidae – finches
*Extant genera present in the fossil record
**Loxia
*Additional prehistoric species of extant genera
**Coccothraustes simeonovi
**Coccothraustes balcanicus
Icteridae – grackles and New World orioles
*Pandanaris
*Pyelorhamphus
*Extant genera present in the fossil record
**Euphagus
Cardinalidae – cardinals
*Placement unresolved
**Passerina sp.
Emberizidae – buntings and New World sparrows
*Pampaemberiza
*Extant genera present in the fossil record
**Ammodramus – including Palaeostruthus
*Additional prehistoric species of extant genera
**Pipilo angelensis
Avialans ''[incertae sedis]''

These fossil taxa cannot be assigned to any major group with reasonable certainty. The "proto-birds" above are of some indeterminate basal position in the entire avialan radiation, but known from such diagnostic material that their relationships at the family level are known. In contrast, the taxa here have a hypodigm that is usually just sufficient for giving them a valid scientific name, but not for phylogenetic purposes beyond classing them as pygostylians or more modern birds. Some, however, are known from such fragmentary remains that the possibility that they are non-avian "reptiles" such as dinosaurs cannot be ruled out at present.

†"Ichthyornis" minusculus Nesov 1990 – enantiornithine?
†Qinornis paleocenica Xue 1995 – enantiornithine? neornithine?
Footnotes

Popular movies

The Hunger Games (film) - 2012 American dystopian action thriller science fiction-adventure film directed by Gary Ross and based on Suzanne Collins’s 2008 novel of the same name. It is the first insta...
untitled Captain Marvel sequel - part of Marvel Cinematic Universe....
Killers of the Flower Moon (film project) - Killers of the Flower Moon - film project in United States of America. It was presented as drama, detective fiction, thriller. The film project starred Leonardo Dicaprio, Robert De Niro. Director of...
Five Nights at Freddy's (film) - Five Nights at Freddy's - film published in 2017 in United States of America. Scenarist of the film - Scott Cawthon....

Popular books

Book of Revelation - The Book of Revelation is the final book of the New Testament, and consequently is also the final book of the Christian Bible. Its title is derived from the first word of the Koine Greek text: apok...
Book of Genesis - account of the creation of the world, the early history of humanity, Israel's ancestors and the origins...
Gospel of Matthew - The Gospel According to Matthew is the first book of the New Testament and one of the three synoptic gospels. It tells how Israel's Messiah, rejected and executed in Israel, pronounces judgement on ...
Michelin Guide - Michelin Guides are a series of guide books published by the French tyre company Michelin for more than a century. The term normally refers to the annually published Michelin Red Guide , the oldest...
Psalms - The Book of Psalms , commonly referred to simply as Psalms , the Psalter or "the Psalms", is the first book of the Ketuvim , the third section of the Hebrew Bible, and thus a book of th...
Ecclesiastes - Ecclesiastes is one of 24 books of the Tanakh , where it is classified as one of the Ketuvim . Originally written c. 450–200 BCE, it is also among the canonical Wisdom literature of the Old Tes...
The 48 Laws of Power - non-fiction book by American author Robert Greene. The book...

Popular television series

The Crown (TV series) - historical drama web television series about the reign of Queen Elizabeth II, created and principally written by Peter Morgan, and produced by Left Bank Pictures and Sony Pictures Tel...
Friends - American sitcom television series, created by David Crane and Marta Kauffman, which aired on NBC from September 22, 1994, to May 6, 2004, lasting ten seasons. With an ensemble cast sta...
Young Sheldon - spin-off prequel to The Big Bang Theory and begins with the character Sheldon...
Modern Family - American television mockumentary family sitcom created by Christopher Lloyd and Steven Levitan for the American Broadcasting Company. It ran for eleven seasons, from September 23...
Loki (TV series) - upcoming American web television miniseries created for Disney+ by Michael Waldron, based on the Marvel Comics character of the same name. It is set in the Marvel Cinematic Universe, shar...
Game of Thrones - American fantasy drama television series created by David Benioff and D. B. Weiss for HBO. It...
Shameless (American TV series) - American comedy-drama television series developed by John Wells which debuted on Showtime on January 9, 2011. It...

List of fossil bird genera

Taxonomic list of fossil prehistoric birds

Basal [Avialae] (extinct)

Basal [Pygostylia] (extinct)

[Enantiornithes] (extinct)

Basal [Euornithes] (extinct)

Basal [Ornithurae] (extinct)

[Neornithes]

[Struthioniformes]

[Casuariiformes]

[Rheiformes]

†[Dinornithiformes]

[Apterygiformes]

†[Lithornithiformes] Houde 1988

[Tinamiformes]

[Vegaviiformes]

[Anseriformes]

[Galliformes]

[Charadriiformes]

†[Gastornithiformes] (extinct)

[Gruiformes]

[Eurypygiformes]

[Cariamiformes]

[Otidiformes]

[Phoenicopteriformes]

[Podicipediformes]

[Phaethontiformes]

[Ciconiiformes]

[Pelecaniformes]

[Procellariiformes]

[Gaviiformes]

[Sphenisciformes]

[Pterocliformes]

[Columbiformes]

[Psittaciformes]

[Opisthocomiformes]

[Musophagiformes]

[Cuculiformes]

[Accipitriformes]

[Falconiformes]

[Steatornithiformes]

[Podargiformes]

[Caprimulgiformes]

[Aegotheliformes]

[Apodiformes]

[Coliiformes]

[Strigiformes]

[Coraciiformes]

[Bucerotiformes]

[Trogoniformes]

[Piciformes]

[Passeriformes]

Avialans ''[incertae sedis]''

Footnotes