Lark


Larks are passerine birds of the family Alaudidae. Larks have a cosmopolitan distribution with the largest number of species occurring in Africa. Only a single species, the horned lark, occurs in North America, and only Horsfield's bush lark occurs in Australia. Habitats vary widely, but many species live in dry regions. When the word "lark" is used without specification, it often refers to the Eurasian skylark .

Taxonomy and systematics

The family Alaudidae was introduced in 1825 by the Irish zoologist Nicholas Aylward Vigors as a subfamily Alaudina of the finch family Fringillidae. Larks are a well-defined family, partly because of the shape of their. They have multiple scutes on the hind side of their tarsi, rather than the single plate found in most songbirds. They also lack a pessulus, the bony central structure in the syrinx of songbirds. They were long placed at or near the beginning of the songbirds or oscines, just after the suboscines and before the swallows, for example in the American Ornithologists' Union's first check-list. Some authorities, such as the British Ornithologists' Union and the Handbook of the Birds of the World, adhere to that placement. However, many other classifications follow the Sibley-Ahlquist taxonomy in placing the larks in a large oscine subgroup Passerida. For instance, the American Ornithologists' Union places larks just after the crows, shrikes, and vireos. At a finer level of detail, some now place the larks at the beginning of a superfamily Sylvioidea with the swallows, various "Old World warbler" and "babbler" groups, and others. Molecular phylogenetic studies have shown that within the Sylvioidea the larks form a sister clade to the family Panuridae which contains a single species, the bearded reedling. The phylogeny of larks was reviewed in 2013, leading to the recognition of the arrangement below.

Extant genera

The family Alaudidae contains 98 extant species which are divided into 21 genera: For more detail, see list of lark species.
ImageGenusLiving Species
Alaemon Keyserling & Blasius, 1840
  • Greater hoopoe-lark
  • Lesser hoopoe-lark
Chersomanes Cabanis, 1851
  • Beesley's lark
  • Spike-heeled lark
  • Ammomanopsis Bianchi, 1905
  • Gray's lark
  • Certhilauda Swainson, 1827
  • Short-clawed lark
  • Karoo long-billed lark
  • Benguela long-billed lark
  • Eastern long-billed lark
  • Cape long-billed lark
  • Agulhas long-billed lark
  • Pinarocorys Shelley, 1902
  • Dusky lark
  • Rufous-rumped lark
  • Ramphocoris Bonaparte, 1850
  • thick-billed lark
  • Ammomanes Cabanis, 1851
  • Desert lark
  • Bar-tailed lark
  • Rufous-tailed lark
  • Eremopterix Kaup, 1836
  • Black-eared sparrow-lark
  • Madagascan lark
  • Black-crowned sparrow-lark
  • Chestnut-backed sparrow-lark
  • Ashy-crowned sparrow-lark
  • Chestnut-headed sparrow-lark
  • Grey-backed sparrow-lark
  • Fischer's sparrow-lark
  • Calendulauda Blyth, 1855
  • Sabota lark
  • Pink-breasted lark
  • Foxy lark
  • Fawn-coloured lark
  • Karoo lark
  • Red lark
  • Dune lark
  • Barlow's lark
  • Heteromirafra Grant, 1913
  • Rudd's lark
  • Archer's lark Clarke, 1920
  • Mirafra Horsfield, 1821
  • Eastern clapper lark
  • Cape clapper lark
  • Red-winged lark
  • Rufous-naped lark
  • Flappet lark
  • Angolan lark
  • Williams's lark
  • Monotonous lark
  • Melodious lark
  • Horsfield's bush lark
  • Singing bush lark
  • Burmese bush lark
  • Bengal bush lark
  • Indochinese bush lark
  • Indian bush lark
  • Jerdon's bush lark
  • Gillett's lark
  • Rusty bush lark
  • Collared lark
  • Ash's lark
  • Somali lark
  • Friedmann's lark
  • Kordofan lark
  • White-tailed lark
  • Lullula Kaup, 1829
  • Woodlark
  • Spizocorys Sundevall, 1872
  • Obbia lark
  • Sclater's lark
  • Stark's lark
  • Short-tailed lark
  • Masked lark
  • Botha's lark
  • Pink-billed lark
  • Alauda Linnaeus, 1758
  • White-winged lark
  • Raso lark
  • Oriental skylark
  • Eurasian skylark
  • Galerida Linnaeus, 1758
  • Sykes's lark
  • Sun lark
  • Large-billed lark
  • Thekla lark
  • Crested lark
  • Malabar lark
  • Maghreb lark
  • Eremophila F. Boie, 1828
  • Horned lark
  • Temminck's lark
  • Calandrella Kaup, 1829
  • Hume's short-toed lark
  • Mongolian short-toed lark
  • Blanford's lark
  • Rufous-capped lark
  • Red-capped lark
  • Greater short-toed lark
  • Melanocorypha F. Boie, 1828
  • Bimaculated lark
  • Calandra lark
  • Black lark
  • Mongolian lark
  • Tibetan lark
  • Chersophilus Sharpe, 1890
  • Dupont's lark
  • Eremalauda WL Sclater, 1926
  • Dunn's lark
  • Alaudala Horsfield & Moore, 1858
  • Athi short-toed lark
  • Asian short-toed lark
  • Somali short-toed lark
  • Lesser short-toed lark
  • Sand lark
  • Extinct genera

  • Genus Eremarida

    Description

  • Larks, which are part of the family Alaudidae, are small- to medium-sized birds, in length and in mass.
    Like many ground birds, most lark species have long hind claws, which are thought to provide stability while standing. Most have streaked brown plumage, some boldly marked with black or white. Their dull appearance camouflages them on the ground, especially when on the nest. They feed on insects and seeds; though adults of most species eat seeds primarily, all species feed their young insects for at least the first week after hatching. Many species dig with their bills to uncover food. Some larks have heavy bills for cracking seeds open, while others have long, down-curved bills, which are especially suitable for digging.
    Larks are the only passerines that lose all their feathers in their first moult. This may result from the poor quality of the chicks' feathers, which in turn may result from the benefits to the parents of switching the young to a lower-quality diet, which requires less work from the parents.
    In many respects, including long tertial feathers, larks resemble other ground birds such as pipits. However, in larks the tarsus has only one set of scales on the rear surface, which is rounded. Pipits and all other songbirds have two plates of scales on the rear surface, which meet at a protruding rear edge.

    Calls and song

    Larks have more elaborate calls than most birds, and often extravagant songs given in display flight. These melodious sounds, combined with a willingness to expand into anthropogenic habitats — as long as these are not too intensively managed — have ensured larks a prominent place in literature and music, especially the Eurasian skylark in northern Europe and the crested lark and calandra lark in southern Europe.

    Behaviour

    Breeding

    Male larks use song flights to defend their breeding territory and attract a mate. Most species build nests on the ground, usually cups of dead grass, but in some species the nests are more complicated and partly domed. A few desert species nest very low in bushes, perhaps so circulating air can cool the nest. Larks' eggs are usually speckled. The size of the clutch is very variable and ranges from the single egg laid by Sclater's lark up to 6-8 eggs laid by the calandra lark and the black lark. Larks incubate for 11 to 16 days.

    In culture

    Larks as food

    Larks, commonly consumed with bones intact, have historically been considered wholesome, delicate, and light game. They can be used in a number of dishes; for example, they can be stewed, broiled, or used as filling in a meat pie. Lark's tongues were particularly highly valued. In modern times, shrinking habitats made lark meat rare and hard to come by, though it can still be found in restaurants in Italy and elsewhere in southern Europe.

    Symbolism

    The lark in mythology and literature stands for daybreak, as in Chaucer's "The Knight's Tale", "the bisy larke, messager of day", and Shakespeare's Sonnet 29, "the lark at break of day arising / From sullen earth, sings hymns at heaven's gate". The lark is also associated with "lovers and lovers' observance" and with "church services", and often those the meanings of daybreak and religious reference are combined (in Blake's Visions of the Daughters of Albion, into a "spiritual daybreak" to signify "passage from Earth to Heaven and from Heaven to Earth". In Renaissance painters such as Domenico Ghirlandaio the lark symbolizes Christ, in reference to John 16:16.

    Pet

    Traditionally larks are kept as pets in China. In Beijing, larks are taught to mimic the voice of other songbirds and animals. It is an old-fashioned habit of the Beijingers to teach their larks 13 kinds of sounds in a strict order. The larks that can sing the full 13 sounds in the correct order are highly valued, while any disruption in the songs will decrease its value significantly.