's assistant Irene O'Dare meets wealthy Don Marshall while she is measuring chairs for Mrs. Herman Vincent at her Long Island estate. Charmed by the young girl, Don anonymously purchases Madame Lucy's, an exclusive Manhattan boutique, and instructs newly hired manager Mr. Smith to offer Irene a job as a model. She soon catches the eye of socialite Bob Vincent, whose mother is hosting a ball at the family mansion. In order to promote Madame Lucy's dress line, Mr. Smith arranges for his models to be invited to the soiree. Irene accidentally ruins the gown she was given to wear and substitutes a quaint blue dress belonging to her mother, and it creates a sensation. Irene is mistaken for the niece of Ireland's Lady O'Dare and, in order to publicize his collection, Mr. Smith decides to exploit the error and moves Irene into a Park Avenue apartment. Dressed in furs and draped with diamonds while escorted around town by Bob, Irene's appearance prompts gossip columnist Biffy Webster to suggest she is a kept woman. Outraged, Irene demands Madame Lucy protect her reputation by revealing the truth, only to discover Don is the owner of the shop. Irene agrees to marry Bob, but on the night before the wedding, Bob confesses he still loves former sweetheart Eleanor Worth, and Irene realizes she loves Don. The couple decides to make things right by reuniting with their rightful partners.
For nearly two decades following its original 1919 production at the Vanderbilt Theatre, Irene — with a total run of 675 performances — held the record for the longest-running show in Broadway theatre history. In addition to the 1926 silent filmIrene, the musical also was adapted for a June 1936Lux Radio Theatre production starring Jeanette MacDonald and Regis Toomey. The film was shot in black and white with the exception of the sequence set at Mrs. Vincent's society ball, featuring the musical number "Alice Blue Gown," which was filmed in Technicolor. The scene is replayed as an installment of Rex Gordon's Moviebone News, a spoof of the Movietone News shorts that were popular at the time. Also seen in the pseudo-news segment are the Dandridge Sisters. Making cameo appearances in the film are Johnny Long and His Orchestra and The Rockettes. This version downplays the "Madame Lucy" character, with the other versions of Irene presenting "Lucy" as actually a very campy gay man.