Interior Low Plateaus
The Interior Low Plateaus are a physiographic region in eastern United States. It consists of a diverse landscape that extends from north Alabama across central Tennessee and Kentucky into southern Illinois, Indiana, and Ohio. Its natural communities are a matrix of temperate forests, woodlands, and prairies.
Setting
This is a region of rolling plains and eroded plateaus, with a Humid subtropical climate in the south and Hot-summer Humid continental climate in the north. It is notable for its extensive karst limestone, which comprise the caves at Mammoth Cave National Park.This region includes a portion of what the U.S. Forest service calls the "Central Hardwood Forest".
The region extends from southern Ohio, Indiana and Illinois through Tennessee and central Kentucky and into northern Alabama. Nearly 65% of Kentucky's territory is within the Interior Low Pleateaus region, which encompasses an area from the western Appalachian Plateaus to the Tennessee River. It includes the Kentucky Bluegrass and Knobs regions, the Highland Rim and the Shawnee Hills.
Geology
The underlying bedrock of the Interior Low Plateaus consists of sedimentary rocks such as limestone, sandstone, and shale. These date from the Ordovician period in the Nashville Basin and Bluegrass region, up to the Carboniferous Period in the Shawnee Hills. The Interior Low Plateaus lie at the southern edge of the glacial boundary. Unlike the till plain to the north, the underlying bedrock is generally close to the surface, and the topography of an area depends on how resistant the underlying bedrock is to erosion. More resistant sandstones have resulted in hillier areas such as the Norman Upland and Crawford Upland in Southern Indiana, while softer limestones have already eroded down to gently rolling plains, like the Scottsburg Lowlands and Wabash Lowlands of southeast and southwest Indiana. The hillier parts of the Interior Low Plateaus are not mountain ranges, but dissected plateaus.Natural Communities
The natural communities in this region are a matrix of forest, woodlands, and prairie. Today, much of the open prairie and savanna communities have been lost due to fire suppression and agriculture. However, oak-hickory woodlands remain relatively common, and mesic forest is abundant along riparian areas.Woodlands
Oak-hickory woodland is the most common natural community in the Interior Low Plateaus. It is the dominant natural community for many areas of rolling hills, including the Western Highland Rim, Shawnee Hills, and Outer Bluegrass. These woodlands represent an intermediate state between a forest and a savanna.Common trees in this community include post oak, white oak, southern red oak, black oak, and pignut hickory. In more calcareous areas, common woodland trees include Shumard oak, Chinquapin oak, and Carolina shagbark. In more acidic dry woodlands there are stands of chestnut oak and Virginia pine. In swampy fragipan flats, woodlands consist of willow oak, overcup oak, pin oak, sweetgum, and blackgum. The woodlands in this region contain a rich sun-loving herbaceous layer, with many species of goldenrod, sunflower, and aster.
Prairie and Savanna
The flat surfaces of the Interior Low Plateaus historically contained extensive areas of open prairie. This fire-maintained community was concentrated in the areas of the Mitchell Plain and Pennyroyal Plain with smaller areas of prairie in the Nashville Basin and Highland Rim. Early settlers describe these areas as nearly treeless expanses, containing Greater prairie chicken and herds of Bison. Today, some prairie remains in areas that are managed by controlled burns, or other forms of tree removal such as powerline and roadside clearings.Dry prairies are the most common remnant in this region, due to their unsuitability for agriculture and resistance to succession. They are dominated by Schizachyrium scoparium and Aristida purpurascens. Mesic prairies are rarer, and are dominated by very tall grasses such as Indian grass, big bluestem, and plume grass. Wet prairies are the rarest and most poorly-documented grassland type in this region. The few existing remnants have species such as gamagrass, prairie cordgrass, and many rushes and sedges. Significant prairie remnants now remain at Ft. Campbell and Arnold Air Force Base, as well as in a few small nature preserves. Blackjack oak was reportedly one of the few trees that could occasionally survive the annual wildfires in the prairies.
Forests
Closed-canopy forests in this region are also naturally found in many areas, such as riparian zones, river bottoms, and dissected terrain.Mesic forests are often found in narrow stream gorges. These forests have a particularly rich diversity in spring wildflowers. They are dominated by sugar maple, beech, basswood, northern red oak, and tuliptree. In the Eastern Highland Rim, at the edge of the Cumberland Plateau, these forests develop a more Appalachian character, and contain species such as eastern hemlock, yellow buckeye, and cucumber magnolia. Acidic sandstone cove forests in the Shawnee Hills also contain more typically Appalachian species such as yellow birch and umbrella magnolia.
Floodplain forests in the Interior Low Plateau remain relatively common, although many areas have been converted to agriculture. These forests are dominated by sugarberry, boxelder, silver maple, and sycamore.
Forests were particularly common in the uplands of the Inner Bluegrass region of Kentucky, which has naturally mesic conditions. These forests were dominated by sugar maple, bitternut hickory, Ohio buckleye, blue ash, and chinquapin oak. This community is now quite rare in the Inner Bluegrass due to logging by early European settlers. Its destruction was so rapid and complete, that mid-20th century ecologists mistook the successional state of the Bluegrass as its natural condition. Sun-loving species such as hackberry, black cherry, black walnut, and white ash now dominate much of the Bluegrass region.
Barrens and Glades
Barrens are open treeless areas on slopes, often with shallow soil. These are found sporadically in the Outer Bluegrass, Inner Nashville Basin, Shawnee Hills, and in parts of the Highland Rim. It is unclear whether these communities rely on fire for their openness, or are maintained by soil conditions alone. They are usually dominated by short grasses such as little bluestem, and large showy forbs such as prairie dock and blazing star. Barrens are particularity abundant and well-preserved in the area of Adams County, Ohio, which includes Adams Lake Prairie, Lynx Prairie, Chaparral Prairie, and others.Glades are areas of flat rock exposures of bedrock. In the Nashville Basin, glades are most often found in areas that were a matrix of barren and woodland communities. Although historically with a more open grassland border, many are now surrounded by dense thickets of redcedar and redbud. Glades are most common the Nashville Basin, although there are rare examples found in the Mitchell Plain, Pennyroyal Plain and Outer Bluegrass. Glades have a flora adapted to extreme dry conditions in the summer, and standing water in the spring. The glades of the Nashville Basin and Moulton Valley are considered a center endemism due to their large number of restricted species. Characteristic species of the Nashville Basin glades include Nashville glade cress, limestone fame-flower, Nashville breadroot, Gattinger's prairie clover, and Tennessee milk-vetch. Glades in other regions contain less endemism, and are characterized by species such as widow's-cross stonecrop, limestone skullcap, glade violet, one-flower gladecress, and poverty dropseed. Glade remnants are common and well-preserved due to their undesirability for agriculture, and their largely edaphic nature. Many examples exist in Cedars of Lebanon State Park, as well as numerous other natural areas.
Wetlands
The Interior Low Plateau has an abundance of sinkhole ponds in the karst regions of the Mitchell Plain, Pennyroyal Plain, and Highland Rim. These isolated communities were historically found within a matrix of wet prairie. These communities are often covered in swamp forest, and are dominated by swamp cottonwood and red maple. High-quality herbaceous examples have species with coastal plain affinities. Many of these communities have been destroyed due to drainage for agriculture.Emergent shrub sloughs are found along the bottoms of major rivers, such as the Cumberland River and Tennessee River. These are dominated by buttonbush, and along the Tennessee River contain bald-cypress. These sloughs were historically widespread, but dam construction and agricultural activity have destroyed most of them.
A rare wetland type in this region is the calcareous seepage fen. These are found sparingly in the Highland Rim, and are dominated by grass of Parnassus and contain the federally-endangered Tennessee yellow-eyed grass. Calcareous seeps are also found near glades in the Inner Nashville Basin. These communities often have extensive stands of sunnybells and other rare species. Calcareous seeps and swamp forest are also found sparingly in the Bluegrass region. These contain trees such as green ash and swamp white oak, along with many herbaceous plants that are considered regionally rare.
Fauna
This region contains the richest diversity of freshwater animals in North America. In particular, the Duck River in Tennessee is considered one of the top three most biologically rich rivers in the world, containing over 150 species of fish, 60 species of freshwater mussels, and 22 species of aquatic snails. The Green River in this region also harbors diversity of a similar scale. This high amount of freshwater mussel diversity is due to the limestone bedrock underlying much of the region. The limestone creates calcareous water that acts as a buffer against acidity, which helps the mussels in shell-building.Birds of the woodlands include vireos and tanagers while mammals include Eastern gray squirrels, chipmunks, raccoons and opossums.
Threats
The dominance of oak in this part of the pre-Columbian savannas of North America was due to frequent fires. The fire suppression policies since the 1930s have been a significant forest disturbance.Today there is very little intact habitat in this ecoregion, with a reduction of bottomland hardwood forests by 70–95%, and only 0.02 percent of the original oak savannas remain. Although much of the area is forested, these forests tend to be highly fragmented and significantly altered by development, agriculture, and fire suppression. The forests are dominated primarily by oak and hickory species, but succession has filled the understory with maples and yellow poplar and blocking oak regeneration. In some areas, habitat is threatened by urbanization and invasive species such as non-native privet, honeysuckle, garlic mustard and kudzu.