Haminoeidae
Haminoeidae, commonly known as the haminoeid bubble snail family, is a taxonomic family of sea snails, marine opisthobranch gastropod mollusks in the superfamily Haminoeoidea.
The name of this family has long been controversial, and used to be Atyidae or Atydidae. Another, but incorrect, spelling was Haminaeidae.
A number of genera have been proposed for this family, but the species are hard to identify by looking only at the external characteristics. Until the internal anatomy of 'wet' specimens has been fully described, the status of many of the genera listed here is uncertain.
Distribution
These bubble snails occur in all warm or temperate seas.Habitat
These are sand dwellers or they live on muddy bottoms, in bays, estuaries, and close to the shore in tidepools.Description of the live animal
These are colorful snails, that can partially take the color of the sea floor.Their large cephalic shield is rounded at the front, but deeply lobed behind. The mantle protrudes behind the shell. The shell is partially or completely enfolded by lateral parapodial lobes.
Shell description
Their shell varies in size according to the species, from 3 mm to 30 mm.The shell is ovoid, thin and translucent. It may be smooth or have spiral grooves. The umbilical apex is sunken or enclosed and no longer visible. Large body whorl with fine spiral striations. Smooth columella. The thin outer lip of the aperture extends beyond the apex of the shell and is thus longer than the body whorl. The aperture narrows posteriorly and is wider anteriorly.
Ecology
These snails are herbivorous. Their diet consists of various kinds of green algae. They can survive in brackish water.They are hermophroditic. Their eggs are deposited in round or oval jellylike strings, attached to eelgrass or sand.
Atys naucum and Atys cylindricum are known to show biological fluorescence.
Genera and species
Genera within the family Haminoeidae include:- Aliculastrum
- Atys Montfort, 1810
- Austrocylichna Burn, 1974
- Bullacta Bergh, 1901 - with the only species: Bullacta exarata
- Cylichnatys Kuroda & Habe, 1952
- Cylichnium W. H. Dall, 1908
- *Cylichnium africanum
- *Cylichnium domitum Dall, 1908 - Distribution: Caribbean
- *Cylichnium matsumotoi - Distribution: Japan
- Diniatys Iredale, 1936
- Haloa
- Haminoea Turton & Kingston in Carrington, 1830
- Liloa Pilsbry, 1921
- Limulatys Iredale, 1936
- Micratys Habe, 1952
- MimatysHabe, 1952
- Mnestia H. Adams & A. Adams, 1854
- Nipponatys Habe, 1952
- Phanerophthalmus A. Adams, 1850
- Sericohaminoea
- Smaragdinella A. Adams, 1848
- Sphaeratys
- Ventomnestia Iredale, 1936
- Weinkauffia Monterosato, 1884
- Atyscaphander Annandale, 1924: synonym of Bullacta Bergh, 1901
- Dinia H. Adams & A. Adams, 1854: synonym of Diniatys Iredale, 1936
- Haminea Gray, 1847: synonym of Haminoea Turton & Kingston in Carrington, 1830
- Sinohaminea Tchang, 1933: synonym of Bullacta Bergh, 1901
- Genus Diniatys
- *Diniatys dentifer A. Adams, 1850 - Distribution: Indo-Pacific, Length : 10 mm, Description : this herbivore is found on the bluegreen algae Lyngbya majuscula, Schizothrix and Hormothamnion. There is a pointed projection on the columella. The color varies between green and various shades of brown. The two black eyes are on the back of the cephalic shield.
- *Diniatys monodonta A. Adams, 1850
- **Distribution: Japan
- Genus Haloa
- *Haloa binotata
- **Distribution : Indo Pacific
- *Haloa constricta A. Adams, 1850
- **Distribution : Japan
- *Haloa crocata W. H. Pease, 1860
- **Distribution : Hawaii
- *Haloa flavescens
- **Distribution : Indo Pacific
- *Haloa fusca W. H. Pease, 1863
- **Distribution : Indo Pacific
- *Haloa japonica H. A. Pilsbry, 1895 Japanese Paper-bubble
- **Distribution : Indo Pacific, Japan
- **Length : 10 mm
- **Description : species with beautiful colors : glassy white background with tiny white spots and dark patches with orange dots.
- *Haloa kawamurai T. Habe, 1950
- **Distribution : Indo Pacific
- *Haloa margaritoides T. Kuroda & T. Habe, 1971
- **Distribution : Japan
- **Length : 7 mm
- **Description : intertidal among seaweeds
- *Haloa nigripunctata W. H. Pease, 1868
- **Distribution : Japan
- *Haloa rotundata A. Adams, 1850
- **Distribution : Japan
- *Haloa vitrea
- **Distribution : Japan
- *Haloa yamaguchii T. Habe, 1952
- **Distribution : Indo Pacific
- Genus Hamineobulla Habe, 1950
- *Hamineobulla kawamurai Habe, 1950
- **Distribution : Okinawa
- **Length : 6 mm
- **Description : brown animal with short cephalic shield; on the shell there are a few transverse rows with brighter dots
- Genus Liloa Pilsbry, 1921
- *Liloa brevis
- **Distribution : Australia
- **Description : elongate body; parapodial flaps cover only the front of the thin, fragile shell
- *Liloa curta
- **Distribution : Western Pacific
- **Description : elongate body; the color can vary from a few dark spots on a translucent body, to almost completely dark.
- *Liloa incisula Yokoyama, 1928
- **Distribution : Japan
- *Liloa laeta A. A. Gould, 1859
- **Distribution : Indo Pacific
- *Liloa nipponensis Nomura & Hatai, 1940
- **Distribution : Japan
- *Liloa porcellana A. A. Gould, 1859
- **Distribution : Japan
- *Liloa tomaculum H. A. Pilsbry, 1951
- **Distribution : Hawaiian Islands
- *Liloa translucens A. Adams, 1862
- **Distribution : Japan
- Genus Limulatys Iredale, 1936
- *Limulatys constrictus T. Habe, 1952
- **Distribution : Indo Pacific
- **Length : 16 mm
- *Limulatys crassilabris
- **Distribution : S.E. Asia, Thailand
- **Length : 8 mm
- *Limulatys fusiformis T. Habe, 1952
- **Distribution : Japan
- *Limulatys muscarius A. A. Gould, 1859
- **Distribution : Indo Pacific, Philippines
- *Limulatys okamotoi T. Habe, 1952
- **Distribution : Indo Pacific
- *Limulatys ooformis T. Habe, 1964 Egg-shaped Bubble
- **Distribution : Indo Pacific, Philippines, Thailand
- **Length : 20 mm
- *Limulatys okamotoi Habe, 1952
- **Distribution : Philippines
- **Length : 9 mm
- *Limulatys reliquus Iredale, 1936
- **Distribution : New Zealand
- *Limulatys scobriculatus A. Adams, 1862
- **Distribution : Japan
- *Limulatys tortuosus A. Adams, 1850
- **Distribution : Japan
- Genus Micratys Habe, 1952
- *Micratys ovum T. Habe, 1952
- **Distribution : Japan, Philippines
- **Length : 2 mm
- Genus Mimatys Habe, 1952
- *Mimatys fukuokaensis T. Habe, 1952
- **Distribution : Japan, Philippines
- **Length : 2.5 mm
- Genus Nipponatys Kurida & Habe, 1952
- *Nipponatys amakusaensis T. Habe & Kikuchi, 1960
- **Distribution : Japan
- *Nipponatys oshimai T. Habe & Kikuchi, 1960
- *Nipponatys volvulina A. Adams, 1862
- **Distribution : Japan
- Genus Sericohaminoea Habe, 1952
- Genus Sphaeratys F. Nordsieck, 1972
- Genus Ventomnestia
- *Ventomnestia bizona
- *Ventomnestia colorata Iredale, 1936
- *Ventomnestia villica
- **Distribution : Guam
- **Length : 5.5 mm
- **Description : a bubble snail with a great variation in color, from white to brown, but always with a characteristic pattern; heavy shell; radula formula : 2.1.2;
- Genus Weinkauffia A. Adams, 1858
- *Weinkauffia diaphana A. Aradas & Maggiore, 1839
- **Distribution : West Africa
- **Description : has similar gizzard plates as Atys multistriatus
- *Weinkauffia turgidula Forbes, 1844
- **Distribution : Adriatic Sea, Malta, Turkey
- **Length : 5 mm
- **Description : fossils of this species have been found in sediments of Pliocene age in Italy.