Epipactis
Epipactis, or Helleborine, is a genus of terrestrial orchids consisting of approximately 70 species. This genus is abbreviated as Epcts in horticultural trade.
The species occur in temperate and subtropical climates of America, Asia, and Europe. These orchids grow in open spaces in forests, in undergrowth, on calcareous soils and are often found in wet dune-slacks near the sea. The only original American species is Giant Helleborine. One species from Europe, Broad-leaved Helleborine, is invasive in North America. Most species are protected.
Most of these hardy orchids grow in a wet environment, but there are exceptions. The Marsh helleborine is the only European orchid able to survive in a flooded habitat. Epipactis gigantea is a species found in the American west, and into southern Canada, in wet areas and even streams. It can grow to a height of 1 m. However, Epipactis helleborine grows in more diverse habitats, from sheltered sandy beaches to open spaces in deciduous or coniferous forests, on roadsides, in meadows, and on moist soils. It is sometimes called the Weed Orchid.
As characteristic of all orchids, Epipactis spp. are dependent on a mycorrhizal symbiosis. This allows some species to have reduced leaves and need little chlorophyll. Violet Helleborine can even do without chlorophyll. These forms can be recognized by their purple instead of violet flowers.
Their creeping, fleshy rhizomes grow offshoots, from which then emerge the 20–70 cm long stems during the next spring.
There are four to eight alternate, lanceolate leaves, that grow progressively shorter near the top. The margins are entire, the top is acute. Species with less chlorophyll have blue-purple leaves.
Their bilaterally symmetrical colorful flowers grow from a terminal raceme. The three sepals and the two lateral petals are ovate and acuminate. Their color can vary from greenish-white to violet and purple.
The lip is divided in a bowl-shaped hypochile, with the outer surface greenish-white and threaded with dark veins. The wavy, snow-white epichile is fan-shaped.
The ovary is inferior. It produces a dry capsule with countless minute seeds.
Species
- Epipactis africana
- Epipactis albensis
- *Epipactis albensis var. albensis. Rhizome geophyte
- *Epipactis albensis var. fibri. Hemicryptophye or rhizome geophyte
- Epipactis aspromontana - now synonym of Epipactis leptochila subsp. aspromontana Kreutz
- Epipactis atromarginata
- Epipactis atrorubens Besser : Dark Red Helleborine, Royal Helleborine
- *Epipactis atrorubens var. atrorubens
- *Epipactis atrorubens var. atrata A.Waldner & Webernd
- *Epipactis atrorubens subsp. danubialis Ciocârlan & R.Rösler
- *Epipactis atrorubens subsp. spiridonovii Kreutz
- *Epipactis atrorubens subsp. subclausa Kreutz
- *Epipactis atrorubens var. triploidea Kreutz
- Epipactis autumnalis D.Doro
- Epipactis baumanniorum Ströhle
- Epipactis bithynica - now synonym of Epipactis helleborine subsp. bithynica Kreutz
- Epipactis bugacensis
- Epipactis campeadorii
- Epipactis cardina
- Epipactis condensata
- Epipactis cretica
- Epipactis danubialis - now synonym of Epipactis atrorubens subsp. danubialis Ciocârlan & R.Rösler
- Epipactis degenii - now synonym of Epipactis halacsyi subsp. degenii Kreutz
- Epipactis distans Arvet-Touvet
- Epipactis dunensis
- Epipactis duriensis Bernardos, et al. - now synonym of Epipactis tremolsii var. duriensis P.Delforge
- Epipactis exilis P.Delforge - now a synonym of Epipactis persica subsp. exilis Kreutz
- Epipactis flaminia - now synonym of Epipactis greuteri var. flaminia Kreutz
- Epipactis flava.
- Epipactis futakii - now synonym of Epipactis leptochila var. futakii P.Delforge
- Epipactis gigantea : Stream Orchid, Chatterbox, Giant Helleborine.
- Epipactis greuteri H.Baumann & Künkele.
- *Epipactis greuteri var. flaminia Kreutz
- *Epipactis greuteri var. preinensis P.Delforge
- *Epipactis greuteri subsp. preinensis
- Epipactis guegelii
- Epipactis halacsyi Robatsch - now basionym of Epipactis viridiflora subsp. halacsyi
- *Epipactis halacsyi subsp. degenii Kreutz
- Epipactis helleborine Crantz : Broad-leaved Helleborine.
- *Epipactis helleborine subsp. bithynica Kreutz
- *Epipactis helleborine subsp. helleborine.
- *Epipactis helleborine subsp. latina.
- *Epipactis helleborine subsp. leutei Kreutz
- *Epipactis helleborine var. minor R.Engel
- *Epipactis helleborine subsp. molochina Kreutz
- *Epipactis helleborine subsp. neerlandica.
- *Epipactis helleborine subsp. orbicularis synonym of Epipactis distans Arvet-Touvet
- *Epipactis helleborine subsp. schubertiorum Kreutz
- *Epipactis helleborine subsp. transcaucasica. Hemicryptophye or rhizome geophyte
- *Epipactis helleborine subsp. tremolsii Hemicryptophye or rhizome geophyte
- *Epipactis helleborine var. youngiana Kreutz
- Epipactis ioessa Bongiorni, De Vivo, Fori & Romolini
- Epipactis kleinii
- Epipactis komoricensis - now synonym of Epipactis leptochila subsp. komoricensis Kreutz
- Epipactis lapidocampi E.Klein & Laminger
- Epipactis latifolia All., nom. illeg.
- *Epipactis latifolia lus. rosea Erdner - now synonym of Epipactis viridiflora var. rosea Kreutz
- Epipactis leptochila Godfery : Narrow-lipped Helleborine.
- *Epipactis leptochila subsp. aspromontana Kreutz
- *Epipactis leptochila var. cleistogama Kreutz
- *Epipactis leptochila var. dinarica P.Delforge
- *Epipactis leptochila var. futakii P.Delforge
- *Epipactis leptochila subsp. komoricensis Kreutz
- *Epipactis leptochila subsp. leptochila
- *Epipactis leptochila subsp. maestrazgona Kreutz
- *Epipactis leptochila subsp. naousaensis Kreutz
- *Epipactis leptochila subsp. neglecta. Hemicryptophyte or rhizome geophyte
- *Epipactis leptochila var. peitzii P.Delforge
- *Epipactis leptochila subsp. sancta Kreutz
- Epipactis leutei - now synonym of Epipactis helleborine subsp. leutei Kreutz
- Epipactis liestalensis.
- Epipactis lusitanica - now synonym of Epipactis tremolsii subsp. lusitanica Kreutz
- Epipactis maestrazgona P.Delforge & Gévaudan
- Epipactis magnibracteata
- Epipactis mairei
- Epipactis maricae - now a synonym of Epipactis nordeniorum subsp. mecsekensis Kreutz
- Epipactis meridionalis H.Baumann & R.Lorenz
- Epipactis microphylla
- Epipactis molochina P.Delforge
- Epipactis moravica Batousek - now a synonym of Epipactis nordeniorum subsp. moravica Kreutz
- Epipactis muelleri.
- *Epipactis muelleri subsp. cerritae. Hemicryptophye or rhizome geophyte
- *Epipactis muelleri subsp. muelleri Rhizome geophyte
- Epipactis nauosaensis - now a synonym of Epipactis leptochila subsp. naousaensis Kreutz
- Epipactis nordeniorum Robatsch
- *Epipactis nordeniorum subsp. mecsekensis Kreutz
- *Epipactis nordeniorum subsp. moravica Kreutz
- Epipactis ohwii
- Epipactis olympica
- Epipactis palustris : Marsh Helleborine, Marsh Orchid
- Epipactis papillosa
- Epipactis persica Nannf.
- *Epipactis persica subsp. exilis Kreutz
- Epipactis phyllanthes G.E.Sm. : Green-flowered Helleborine.
- *Epipactis phyllanthes subsp. fageticola Kreutz
- *Epipactis phyllanthes var. olarionensis. Hemicryptophye or rhizome geophyte
- *Epipactis phyllanthes var. phyllanthes
- Epipactis placentina.
- Epipactis pollinensis - now synonym of Epipactis viridiflora var. pollinensis Kreutz
- Epipactis pontica
- Epipactis provincialis
- Epipactis pseudopurpurata - Epipactis viridiflora subsp. pseudopurpurata Kreutz
- Epipactis rechingeri
- Epipactis rhodanensis
- Epipactis robatschiana Bartolo, D'Emerico, Pulv., Terrasi & Stuto
- Epipactis royleana
- Epipactis sancta: Lindisfarne helleborine
- Epipactis schubertiorum - now synonym of Epipactis helleborine subsp. schubertiorum Kreutz
- Epipactis spiridonovii - now synonym of Epipactis atrorubens subsp. spiridonovii Kreutz
- Epipactis stellifera
- Epipactis subclausa - now synonym of Epipactis atrorubens subsp. subclausa Kreutz
- Epipactis tallosii
- Epipactis tenii
- Epipactis thessala
- Epipactis thunbergii
- Epipactis tremolsii Pau
- *Epipactis tremolsii subsp. densifolia Kreutz
- *Epipactis tremolsii var. duriensis P.Delforge
- *Epipactis tremolsii subsp. heraclea Kreutz
- *Epipactis tremolsii subsp. lusitanica Kreutz
- *Epipactis tremolsii subsp. turcica Kreutz
- Epipactis troodi
- Epipactis turcica - now synonym of Epipactis tremolsii subsp. turcica Kreutz
- Epipactis turcomanica
- Epipactis ulugurica
- Epipactis veratrifolia : Scarce Marsh Helleborine
- Epipactis viridiflora Krock: Clustered Helleborine, Violet Helleborine
- *Epipactis viridiflora subsp. halacsyi B.Baumann & H.Baumann
- *Epipactis viridiflora subsp. pollinensis B.Baumann & H.Baumann
- *Epipactis viridiflora subsp. pseudopurpurata Kreutz
- *Epipactis viridiflora var. rosea Kreutz
- Epipactis xanthophaea
- Epipactis youngiana : Young's Helleborine - now synonym of Epipactis helleborine var. youngiana Kreutz
Hybrids
- Epipactis × amigoi .
- Epipactis × barlae .
- Epipactis × barreana .
- Epipactis × breinerorum .
- Epipactis × bruxellensis .
- Epipactis × capellonensis .
- Epipactis × cardonneae .
- Epipactis × conquensis .
- Epipactis × gerbaudiorum
- Epipactis × gevaudanii .
- Epipactis × graberi .
- Epipactis × heterogama .
- Epipactis x nicolosii M.P.Grasso & Grillo
- Epipactis × populetorum .
- Epipactis × pupplingensis .
- Epipactis × reinekei .
- Epipactis x robatschii Gévaudan & P.Delforge
- Epipactis × schmalhausenii .
- *Epipactis × schmalhausenii nothosubsp. fleischmannii . Hemicryptophye or rhizome geophyte
- *Epipactis × schmalhausenii nothosubsp. schmalhausenii. Rhizome geophyte
- Epipactis × schulzei.
- Epipactis × soguksuensis.
- Epipactis × stephensonii .
- Epipactis × trikalana .
- Epipactis × vermionensis .