Eogruidae is a family of large, flightless birds that occurred across Eurasia from the Eocene to Pliocene epochs. Related to modern cranes, limpkins and trumpeters, these birds converged strongly with ostriches due to similar speciations to cursoriality, even reducing their number of toes to two in some taxa. It has been suggested that competition from true ostriches has caused the extinction of these birds, though this has never been formally tested and several ostrich taxa do occur in the late Cenozoic of Asia and some species do occur in areas where ostrich fossils have also been found.
Description
Most eogruids are known from rather sparse remains, mostly the tarso-metatarsals and toes. The former are generally slender with a distinct crest along the lateral side of the plantar surface. The trochlea for the second toe shows a progressive reduction along the various taxa, culminating in its utterabsence in Amphipelargus. Other skeletal remains are rare. Wing elements are known in a few taxa; these are generally highly reduced, suggesting that these birds were flightless. The exception appears to be Eogrus itself, which does not show much phalange reduction and hypothetically could still fly, though the rest of the postcranial skeleton is already specialised for cursoriality. '', with respective living relatives; a tinamou and a trumpeter, illustrating the degree of convergent evolution
Classification
Eogruids are almost universally accepted to be Gruiformes within the crane-limpkin-trumpeter line, Gruoidea, the exception being Olson 1985 which declared them to be stem-ostriches, a study whose proposals are not taken seriously; any similarities between eogruiids and ostriches are purely convergent in nature. More derived taxa such as Ergilornis and Amphipelargus were sometimes classified as a separate family, Ergilornithidae, but they are now generally accepted to be a subfamily within Eogruidae. Eogruids are rather similar to Geranoididae, a clade of similar flightless gruiforms from North America and Europe, and both groups have occasionally been classified as sister taxa. Recently geranoidids have been recovered as basal to the rest of Gruoidea, however, while eogruids are sister-taxa to cranes.
Taxonomy
There is some contention in regards to ergilornithine genera, particularly Amphipelargus and Urmiornis, which have assimilated each other's species from study to study. The most recent consensus is that nearly all Neogene eogruid remains belong to Urmiornis, while a few western Eurasian taxa can be referred to as Amphipelargus.
