Dunkleosteidae


Dunkleosteidae is an extinct family of arthrodire placoderms. The gigantic apex predator Dunkleosteus terrelli is the best known member of this group. While they were previously thought to be close relatives of the genus Dinichthys and grouped together in the family Dinichthyidae, more recent studies have shown that the two taxa represent two very distinct clades within Arthrodira. The reappraisal of Kiangyousteus lead to a restructuring of the family, with the inclusions of the benthic, aberrant Heterosteus as the sister taxon of Dunkleosteus, and the Late Emsian Xiangshuiosteus as the sister taxon of Eastmanosteus calliaspis, and the removal of Westralichthys from the family

Genera

''Dunkleosteus''

The type genus, Dunkleosteus, is known from Late Frasnian and Famennian-aged marine strata from Europe, Morocco, and North America. The best known species, D. terrelli, is famous as the "world's first vertebrate apex predator," and is estimated to be up to in length: other species, however, such as D. raveri, are estimated to be in length.

''Eastmanosteus''

Eastmanosteous is a diverse genus of medium to somewhat large predatory arthrodires very similar in anatomy to the species of Dunkleosteus. Eastmanosteus differs from Dunkleosteus in having a unique tubercle-ornamentation on the dermal surfaces of the plates, a distinctively shaped nuchal plate, and sutures that are more zigzagging. The best studied species, E. calliaspis may not be of this genus due to its recently appreciated relationship to the Emsian-aged genus Xiangshuiosteus.

''Golshanichthys''

Fossils of Golshanichthys are found in Frasnian-aged marine strata near Kerman, Iran.

''Kiangyousteus''

This Middle Devonian genus represents the first arthrodire described from China. Fossils are known from the Late Givetian to Early Frasnian-aged Guanwu Formation in Sichuan.

''Xiangshuiosteus''

Xiangshuiosteus was originally described as an arthrodire incertae sedis with anatomical features suggestive of both buchanosteids and coccosteids. With the reappraisal of Kiangyousteus, it is now thought to be a dunkleosteid most closely related to Eastmanosteus calliaspis.