Dactylorhiza
Dactylorhiza is a genus of flowering plants in the orchid family Orchidaceae. Its species are commonly called marsh orchids or spotted orchids,Dactylorhiza were previously classified under Orchis which has two round tubers.
Description
They are hardy tuberous geophytes. In a thickened underground stem, they can store a large amount of water to survive arid conditions. The tuber is flattened and finger-like. The long leaves are lanceolate and, in most species, also speckled. They grow along a rather long stem which reaches a height of. Leaves higher on the stem are shorter than leaves lower on the stem. The inflorescence, compared to the length of the plant, is rather short. It consists of a compact raceme with 25-50 flowers. These develop from axillary buds. The dominant colors are white and all shades of pink to red, sprinkled with darker speckles.Taxonomy
Etymology
The name Dactylorhiza is derived from Greek words δάκτυλος "daktylos" and ρίζα "rhiza", referring to the palmately two- to five-lobed tubers of this genus.Species
Many species in this genus hybridise so readily that species boundaries themselves are vague, with regular name changes and no clear answers. A few species colonise very well onto fresh industrial wastes such as pulverised fuel ash, where vast hybrid swarms can appear for a decade or more, before ecological succession replaces them.- Dactylorhiza alpestris : Alpine Dactylorhiza.
- Dactylorhiza angustata.
- Dactylorhiza aristata : Keyflower.
- *Dactylorhiza aristata var. aristata : Keyflower.
- *Dactylorhiza aristata var. kodiakensis : Kodiak Keyflower.
- Dactylorhiza armeniaca - has become synonym of Dactylorhiza euxina subsp. armeniaca Kreutz
- Dactylorhiza atlantica Kreutz & Vlaciha
- Dactylorhiza baldshuanica.
- Dactylorhiza baltica
- Dactylorhiza baumanniana.
- *Dactylorhiza baumanniana subsp. smolikana H. Baumann & R. Lorenz
- Dactylorhiza bohemica.
- Dactylorhiza cordigera Soó .
- *Dactylorhiza cordigera subsp. bosniaca.
- *Dactylorhiza cordigera subsp. cordigera.
- *Dactylorhiza cordigera var. graeca
- *Dactylorhiza cordigera subsp. pindica H. Baumann & R. Lorenz .
- *Dactylorhiza cordigera var. rhodopeia Presser
- *Dactylorhiza cordigera subsp. siculorum.
- Dactylorhiza ebudensis P. Delforge : Hebridean Marsh Orchid
- Dactylorhiza elata Soó : Stately Dactylorhiza.
- *Dactylorhiza elata subsp. ambigua Kreutz
- *Dactylorhiza elata subsp. brennensis.
- *Dactylorhiza elata subsp. elata.
- *Dactylorhiza elata subsp. mauritanica B.Baumann & H. Baumann
- *Dactylorhiza elata subsp. sesquipedalis.
- Dactylorhiza euxina Czerep.
- *Dactylorhiza euxina subsp. armeniaca Kreutz
- Dactylorhiza flavescens.
- Dactylorhiza foliosa : Richly leaved Dactylorhiza.
- Dactylorhiza fuchsii Soó : Common Spotted Orchid, Fuch's Dactylorhiza .
- *Dactylorhiza fuchsii subsp. carpatica Kreutz
- *Dactylorhiza fuchsii subsp. fuchsii.
- *Dactylorhiza fuchsii subsp. hebridensis.
- *Dactylorhiza fuchsii subsp. meyeri Kulikov & E.G.Philippov
- *Dactylorhiza fuchsii subsp. okellyi.
- *Dactylorhiza fuchsii subsp. psychrophila.
- *Dactylorhiza fuchsii var. sooana Kreutz
- *Dactylorhiza fuchsii var. sudetica H.Baumann
- Dactylorhiza gervasiana.
- Dactylorhiza graeca - has become synonym of Dactylorhiza cordigera var. graeca
- Dactylorhiza graggeriana.
- Dactylorhiza hatagirea.
- Dactylorhiza iberica.
- Dactylorhiza ilgazica - now synonym of Dactylorhiza urvilleana subsp. ilgazica Kreutz
- Dactylorhiza incarnata Soó : Early Marsh Orchid
- *Dactylorhiza incarnata var. baumgartneriana P.Delforge
- *Dactylorhiza incarnata subsp. coccinea
- *Dactylorhiza incarnata subsp. cruenta.
- *Dactylorhiza incarnata subsp. gemmana.
- *Dactylorhiza incarnata subsp. incarnata.
- *Dactylorhiza incarnata nothosubsp. krylovii.
- *Dactylorhiza incarnata subsp. lobelii.
- *Dactylorhiza incarnata subsp. ochroleuca.
- *Dactylorhiza incarnata subsp. pulchella.
- *Dactylorhiza incarnata nothosubsp. versicolor
- Dactylorhiza insularis : Island Dactylorhiza.
- Dactylorhiza kafiriana.
- Dactylorhiza kalopissii E.Nelson.
- *Dactylorhiza kalopissii subsp. macedonica Kreutz
- *Dactylorhiza kalopissii subsp. pythagorae Kreutz
- Dactylorhiza kulikalonica.
- Dactylorhiza lapponica Soó.
- *Dactylorhiza lapponica subsp. angustata Kreutz
- *Dactylorhiza lapponica subsp. rhaetica H. Baumann & R. Lorenz
- Dactylorhiza libanotica
- Dactylorhiza longifolia.
- Dactylorhiza macedonica - now a synonym of Dactylorhiza kalopissii subsp. macedonica Kreutz
- Dactylorhiza maculata Soó : Heath Spotted Orchid, Moorland Spotted Orchid.
- *Dactylorhiza maculata subsp. battandieri.
- *Dactylorhiza maculata subsp. caramulensis.
- *Dactylorhiza maculata subsp. elodes.
- *Dactylorhiza maculata subsp. ericetorum : Heath Spotted Orchid.
- *Dactylorhiza maculata subsp. islandica.
- *Dactylorhiza maculata subsp. kolaensis Kreutz
- *Dactylorhiza maculata subsp. maculata.
- *Dactylorhiza maculata subsp. maurusia
- *Dactylorhiza maculata subsp. podesta.
- *Dactylorhiza maculata subsp. rhoumensis
- *Dactylorhiza maculata subsp. savogiensis Kreutz
- *Dactylorhiza maculata subsp. schurii.
- *Dactylorhiza maculata subsp. sennia Kreutz
- *Dactylorhiza maculata subsp. transsilvanica.
- Dactylorhiza magna.
- Dactylorhiza majalis P.F.Hunt & Summerh. : Broad-leaved Marsh Orchid, Western Marsh Orchid, Fan Orchid, Common Marsh Orchid.
- *Dactylorhiza majalis var. brevifolia Kreutz
- *Dactylorhiza majalis subsp. calcifugiens
- *Dactylorhiza majalis subsp. majalis.
- *Dactylorhiza majalis subsp. occidentalis
- *Dactylorhiza majalis subsp. parvimajalis Kreutz
- *Dactylorhiza majalis subsp. sphagnicola
- *Dactylorhiza majalis subsp. turfosa - has become a synonym of Dactylorhiza traunsteineri subsp. turfosa Kreutz
- Dactylorhiza markusii : Markus' Dactylorhiza.
- Dactylorhiza nieschalkiorum.
- Dactylorhiza occidentalis P. Delforge : Irish Marsh Orchid
- Dactylorhiza osmanica.
- *Dactylorhiza osmanica var. anatolica.
- *Dactylorhiza osmanica var. osmanica.
- Dactylorhiza pindica.
- Dactylorhiza praetermissa : Leopard Marsh Orchid, Southern Marsh Orchid
- Dactylorhiza purpurella Soó : Northern Marsh Orchid.
- *Dactylorhiza purpurella var. maculosa
- *Dactylorhiza purpurella var. purpurella
- *Dactylorhiza purpurella var. cambrensis R.M.Bateman & Denholm 2005
- Dactylorhiza pythagorae - now a synonym of Dactylorhiza kalopissii subsp. pythagorae Kreutz
- Dactylorhiza romana : Roman Dactylorhiza
- Dactylorhiza russowii Holub
- Dactylorhiza saccifera Soó : Sack-carrying Dactylorhiza
- *Dactylorhiza saccifera subsp. bithynica Kreutz
- *Dactylorhiza saccifera subsp. gervasiana Kreutz
- Dactylorhiza salina
- Dactylorhiza sambucina : Elder-flowered Orchid.
- Dactylorhiza sudetica
- Dactylorhiza traunsteineri Soó : Narrow-leaved Marsh Orchid, Traunstein's Dactylorhiza.
- *Dactylorhiza traunsteineri subsp. carpatica - has become synonym of Dactylorhiza fuchsii subsp. carpatica Kreutz
- *Dactylorhiza traunsteineri subsp. turfosa Kreutz
- *Dactylorhiza traunsteineri subsp. curvifolia.
- *Dactylorhiza traunsteineri subsp. traunsteineri.
- *Dactylorhiza traunsteineri subsp. wirtgenii Kreutz
- *Dactylorhiza traunsteinerioides Landwehr
- Dactylorhiza umbrosa
- Dactylorhiza urvilleana H.Baumann & Künkele
- *Dactylorhiza urvilleana subsp. bithynica H. Baumann & R. Lorenz
- *Dactylorhiza urvilleana subsp. phoenissa B. Baumann & H. Baumann
- *Dactylorhiza urvilleana subsp. ilgazica Kreutz
- Dactylorhiza viridis : Frog orchid.
- *Dactylorhiza viridis var. virescens
- *Dactylorhiza viridis var. viridis
Hybrids
- Dactylorhiza × abantiana .
- Dactylorhiza × aldenii .
- Dactylorhiza × altobracensis .
- Dactylorhiza × aschersoniana .
- *Dactylorhiza × aschersoniana nothosubsp. aschersoniana.
- *Dactylorhiza × aschersoniana nothosubsp. templinensis .
- *Dactylorhiza × aschersoniana nothovar. uliginosa .
- Dactylorhiza × baicalica .
- Dactylorhiza × balabaniana .
- Dactylorhiza × bayburtiana .
- Dactylorhiza × beckeriana.
- Dactylorhiza × boluiana .
- Dactylorhiza × bourdonii .
- Dactylorhiza × braunii .
- *Dactylorhiza × braunii nothosubsp. braunii.
- *Dactylorhiza × braunii nothosubsp. lilacina .
- *Dactylorhiza × braunii nothosubsp. monticola .
- *Dactylorhiza × braunii nothosubsp. smitakii . te
- Dactylorhiza × breviceras .
- Dactylorhiza × carnea .
- *Dactylorhiza × carnea nothosubsp. ampolai .
- *Dactylorhiza × carnea nothosubsp. carnea.
- *Dactylorhiza × carnea nothosubsp. maculatiformis. .
- Dactylorhiza × claudiopolitana .
- Dactylorhiza × csatoi .
- Dactylorhiza czerniakowskae.
- Dactylorhiza × daunia .
- Dactylorhiza × delamainii .
- Dactylorhiza × dinglensis .
- *Dactylorhiza × dinglensis nothosubsp. dinglensis.
- *Dactylorhiza × dinglensis nothosubsp. robertsii .
- *Dactylorhiza × dinglensis nothosubsp. senayi .
- *Dactylorhiza × dinglensis nothosubsp. townsendiana .
- *Dactylorhiza × dinglensis nothosubsp. vermeuleniana .
- Dactylorhiza × drucei
- Dactylorhiza × dubreuilhii .
- Dactylorhiza × dufftiana .
- Dactylorhiza × dufftii .
- *Dactylorhiza × dufftii nothosubsp. dufftii.
- *Dactylorhiza × dufftii nothosubsp. gotlandica . Tuber geophyte
- *Dactylorhiza × dufftii nothosubsp. stenkyrkae .
- Dactylorhiza × erdingeri .
- Dactylorhiza euxina.
- *Dactylorhiza euxinavar. euxina.
- *Dactylorhiza euxinavar. markowitschii.
- Dactylorhiza × flixensis .
- Dactylorhiza × formosa .
- Dactylorhiza × fourkensis .
- Dactylorhiza × gabretana .
- Dactylorhiza × genevensis .
- Dactylorhiza × godferyana .
- Dactylorhiza × grandis .
- Dactylorhiza × guilhotii .
- Dactylorhiza × guillaumeae .
- Dactylorhiza × gustavssonii .
- Dactylorhiza × hallii .
- *Dactylorhiza × hallii nothosubsp. hallii.
- *Dactylorhiza × hallii nothosubsp. nummiana .
- Dactylorhiza × hochreutinerana .
- Dactylorhiza × insignis .
- Dactylorhiza × ishorica .
- Dactylorhiza × jenensis
- Dactylorhiza × jestrebiensis .
- Dactylorhiza × juennensis .
- Dactylorhiza × katarana .
- Dactylorhiza × kelleriana .
- Dactylorhiza × kerasovinensis .
- Dactylorhiza × kerneriorum .
- * Dactylorhiza × kerneriorum nothosubsp. kerneriorum.
- *Dactylorhiza × kerneriorum nothosubsp. lillsundica .
- *Dactylorhiza × kerneriorum nothosubsp. variablis . *Dactylorhiza × komiensis .
- Dactylorhiza × kopdagiana .
- Dactylorhiza × koutsourana .
- Dactylorhiza × kuuskiae .
- Dactylorhiza × latirella .
- Dactylorhiza × lehmannii .
- Dactylorhiza × megapolitana .
- Dactylorhiza × metsowonensis .
- Dactylorhiza × mixtum .
- Dactylorhiza × mulignensis .
- Dactylorhiza × nevskii .
- Dactylorhiza × ornonensis .
- Dactylorhiza × paridaeniana .
- Dactylorhiza × pontica
- Dactylorhiza × prochazkana .
- Dactylorhiza × renzii .
- Dactylorhiza × rizeana .
- Dactylorhiza × rombucina .
- Dactylorhiza × ruppertii .
- Dactylorhiza × salictina .
- Dactylorhiza × serbica .
- Dactylorhiza × serreana .
- Dactylorhiza × sivasiana .
- Dactylorhiza × sooi .
- Dactylorhiza × souflikensis .
- Dactylorhiza × stagni-novi .
- Dactylorhiza × szaboiana .
- Dactylorhiza × transiens
- *Dactylorhiza × transiens nothosubsp. corylensis
- *Dactylorhiza × transienssubsp. ericetorum. Tuber geophyte
- *Dactylorhiza × transiens nothosubsp. transiens. Tuber geophyte
- Dactylorhiza × turcestanicum .
- Dactylorhiza × vallis-peenae .
- Dactylorhiza × venusta .
- *Dactylorhiza × venusta nothosubsp. hebridella .
- *Dactylorhiza × venusta nothosubsp. venusta
- Dactylorhiza × viridella .
- Dactylorhiza × vitosana .
- Dactylorhiza × vogtiana .
- Dactylorhiza × vorasica .
- Dactylorhiza × weissenbachiana .
- Dactylorhiza × wiefelspuetziana .