There are three villages within the Cuckmere Valley: Litlington, Lullington and Westdean. Archaeological remains nearer the coast at Exceat suggest a further settlement there in former times. The civil parish of Cuckmere Valley was created in 1990 when Litlington, Lullington and Westdean parishes were abolished, with small areas transferred between Litlington and Alfriston, which remained a separate civil parish.
Lullington is the most northerly of the three, about southeast of Lewes. It lies on a shoulder of the South Downs at the point where the River Cuckmere cuts through the downs, on the opposite bank of the river to Alfriston. A medieval village, Lullington Court, was abandoned after the Black Death in the 14th century; its site was to the east of the present Lullington. The church, dedicated to the Good Shepherd, is believed to be the smallest in England, having been rebuilt using only the original chancel after a fire had destroyed the remainder of the building. Services are held only in the summer months, as the building has no electricity.
Exceat
Exceat was an ancient village. The A259 goes by it just east of the River Cuckmere. The name may have come from æc-sceat, an oak grove, or from the Old English for "the place of the Aese", early kings of Kent whilst another etymology is possible from Brittonic wysg meaning water or river and ceat meaning woods, hence "Wooded River" or "Wooded Ridge". After the Norman conquest, the village was given to Robert, Count of Mortain, half brother of William the Conqueror. Both Exceat and the land were given later to Grestein Priory in Wilmington. Until the 14th century, Exceat was an important village with its own church. The village suffered French raids; it suffered many fatalities during the Black Death, as was effectively abandoned by the 15th century. The main features are the Visitors Centre for the nearby Seven Sisters and the Cuckmere Inn Public House across Exceat Bridge over the Cuckmere.
West Dean is located on the edge of Friston Forest. Its church is dedicated to All Saints.
Landmarks
The National Trust owns land on the western side of the river south of the A259. In addition, there are three Sites of Special Scientific Interest within the parish; Seaford to Beachy Head, Wilmington Downs and Lullington Heath. Seaford to Beachy Head is a large site of biological and geological importance. The area contains a wide variety of habitats based on the chalk bedrock, which hosts a wide range of flora and fauna, including some nationally rare species. Wilmington Downs is another area of biological interest; its chalk grassland is habitat for several rare species. Lullington Heath is also of biological interest due to its chalk heath and grassland habitats.
