Cochliopidae Cochliopidae is a family of small freshwater snails with gills and an operculum , aquatic gastropod mollusks. Paludestrina d'Orbigny, 1840 is an archaic synonym, and has been placed on the Official Index of Rejected and Invalid Names by ICZN Opinion 2202 . This family is in the superfamily Truncatelloidea and in the clade Littorinimorpha .Description Cochliopidae snails are characterized by sharp, elongated spire-like structures, and can be found in rivers or brackish water .2005 taxonomy The family Cochliopidae consists of 3 subfamilies :Cochliopinae Tryon , 1866 - synonyms: Mexithaumatinae D. W. Taylor , 1966, Paludiscalinae D. W. Taylor, 1966 Littoridininae Thiele , 1928 Semisalsinae Guiusti & Pezzoli, 1980 - synonym: Heleobiini Bernasconi, 1991Genera Liu et al . have recognized 34 genera with more than 260 species within the subfamily Cochliopinae .Strong et al. have recognized 246 freshwater species within Cochliopidae. Genera within the family Cochliopidae include: subfamily Cochliopinae subfamily Littoridininae Antrobia Hubricht, 1971 - with the only species Antrobia culveri Hubricht, 1971 - Tumbling Creek cavesnail Littoridina Souleyet , 1852Pseudotryonia Hershler, 2001*Pseudotryonia adamantina *Pseudotryonia alamosae *Pseudotryonia brevissima *Pseudotryonia grahamae *Pseudotryonia mica Hershler, Liu & Landye, 2011 *Pseudotryonia pasajae Hershler, Liu & Landye, 2011 subfamily Semisalsinae - there are three genera in the subfamily SemisalsinaeHeleobia Stimpson, 1865Heleobops Thompson, 1968Semisalsa Radoman, 1974 subfamily ? Aroapyrgus H. B. Baker, 1931Balconorbis Hershler & Longley, 1986† Carinulorbis Yen, 1949 Chorrobius Hershler, Liu & Landye, 2011Dyris Conrad, 1871 - it has extant species and also 26 species in Miocene Pebas Formation Emmericiella Pilsbry , 1909Eremopyprgus Hershler, 1999† Feliconcha Wesselingh, Anderson & Kadolsky, 2006 - two species from Miocene of the Pebas Formation *Feliconcha feliconcha Wesselingh, Anderson & Kadolsky, 2006 *Feliconcha reticulata Wesselingh, Anderson & Kadolsky, 2006 Glabertryonia Wesselingh, Anderson & Kadolsky, 2006 - three species *† Glabertryonia glabra Wesselingh, Anderson & Kadolsky, 2006 - from Miocene of the Pebas Formation *† Glabertryonia sp. 1 - from Pliocene of the Las Piedras Formation * Glabertryonia sp. 2 - from Holocene of Surinam, probably extant Juturnia Hershler, Liu & Stockwell, 2002Lithococcus Pilsbry, 1911 Mesobia F. G. Thompson & Hershler, 1991Mexipyrgus Taylor, 1966Minckleyella Hershler, Liu & Landye, 2011 - with the only species Minckleyella balnearis Hershler, Liu & Landye, 2011 Onobops Thompson, 1968 Pyrgophorus Ancey, 1888 Sioliella Haas, 1949 Spurwinkia Davis, Mazurkiewicz & Mandracchia, 1982 Texadina Abbott & Ladd, 1951 Thalassobia Bourguignat in Mabille, 1877 Cladogram A cladogram based on sequences of mitochondrial cytochrome-c oxidase I genes showing phylogenic relations of species within Cochliopidae:
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