Cassidae


The Cassidae are a taxonomic family of medium-sized, large, and sometimes very large sea snails commonly called helmet snails or bonnet snails. These are marine gastropod mollusks in the superfamily Tonnoidea and the clade Littorinimorpha.
About 60 species comprise the family Cassidae; an example is Cypraecassis rufa.

Nomenclature

Despite its incorrect formation, the ICZN has placed the name Cassidae Latreille, 1825 on the official list of family names, therefore avoiding homonymy with Cassididae Stephens, 1831 ; Opinion 1023.

Distribution

Species of this family occur in tropical and temperate seas from the intertidal zone to depths of 100 m, buried in the sand during the day and becoming active at night.

Shell description

Members of this family are shaped rather like bonnets or helmets, as their common name suggests. The shells are large, thick, subglobular with dextrally coiled, sometimes varicose, whorls, and a short spire. The coiling may be trochospiral or convoluted. The shells of many species have great variability, which has led to many misidentifications, resulting in many synonyms.
Many species have a large and solid shield over the parietal body or beside the thick, plicated columella. Many species show blunt knobs and thickened axial ridges, known as varices. The thin, horny operculum is oval in shape and covers a long aperture. The siphonal canal is straight or slightly curved. The outer lip is somewhat thicker at its margin and toothed on the inside.

Anatomy

These snails have a large mantle and a large, muscular foot. Their large head has an extensible snout. The eyes are at the base of the single pair of tentacles.

Feeding habits

Helmet bonnets prey on echinoderms, starting by gripping them using the foot. The snail then makes a hole in the urchin through the combined action of a secretion which is rich in sulfuric acid and by rasping with their radula. The acid secretion is provided by two large proboscis glands.

Taxonomy

In their taxonomy of the Gastropoda of 2005 Bouchet & Rocroi still listed Cassidae as Cassinae Latreille, 1825, a subfamily of Tonnidae Suter, 1913, following in this Riedel. In his "Revision of the deep-water Cassidae" however, Beu raised in 2008 Cassinae to the rank of family.

Genera

Genera within the family include:
;Genera brought into synonymy :
Some helmet shells are carved into cameos, starting in Italy in 1820.