Canthium Canthium is a genus of flowering plants in the family Rubiaceae . They are shrubs and small trees . The leaves are deciduous and the stems are usually thorny .Distribution Canthium species are predominantly found in Southeast Asia , especially in Thailand and the Philippines . A small number of species is found in India , Sri Lanka , and Bangladesh . Only a limited number of species is found on the African continent , especially in Southern and East Africa .Taxonomy Canthium was named by Jean-Baptiste Lamarck in 1785 in Encyclopédie Méthodique . The name is a latinisation of "kantankara", a Malayalam name from Kerala for Canthium coromandelicum . Kantan means "shining" and kara means "a spiny shrub". The biological type for the genus consists of specimens originally described by Jean-Baptiste Lamarck as Canthium parviflorum but this species is now included in Canthium coromandelicum . Canthium is a member of Vanguerieae , a tribe that is monophyletic and easily recognized morphologically , but in which generic boundaries were, for a long time, very unclear. Canthium was especially problematic, and until the 1980s, it was defined broadly and known to be polyphyletic . Psydrax was separated from it in 1985, as was Keetia in 1986. These were followed by Pyrostria and Multidentia in 1987. The subgenus Afrocanthium was raised to generic rank in 2004, followed by Bullockia in 2009 . A few species were transferred to Canthium from Rytigynia and other genera in 2004. The genus was further reduced by the transfer of species to Peponidium and Pyrostria . In 2016, two Canthium species endemic to the Philippines were transferred to a genus of their own, Kanapia . The final circumscription of Canthium will remain in doubt until phylogenetic studies achieve greater resolution for the clade containing Canthium coromandelicum and its closest relatives.Species Canthium aciculatum Ridl. Canthium angustifolium Roxb. Canthium approximatum Korth. Canthium arboreum Vidal Canthium aurantiacum Merr. & L.M.Perry Canthium berberidifolium Geddes Canthium bipinnatum Merr. Canthium brunneum Merr. Canthium calvum Craib Canthium cambodianum Pit. Canthium campanulatum Thwaites Canthium cavaleriei H.Lév. Canthium ciliatum Kuntze Canthium coffeoides Pierre ex Pit. Canthium confertum Korth. Canthium congestiflorum Ridl. Canthium cordatum Dillwyn Canthium coromandelicum Alston Canthium culionense Merr. Canthium depressinerve Ridl. Canthium ellipticum Merr. Canthium elmeri Merr. Canthium fenicis Merr. Canthium ferrugineum Craib Canthium filipendulum Pierre ex Pit. Canthium fraternum Miq. Canthium glandulosum Merr. Canthium glaucum Hiern Canthium gracilipes Kurz Canthium hainanense Lantz Canthium hirtellum Ridl. Canthium hispidonervosum C.M.Evrard Canthium horridulum Craib Canthium horridum Blume Canthium inerme Kuntze Canthium korthalsianum Miq. Canthium kuntzeanum Bridson Canthium laeve Teijsm. & Binn . Canthium lasianthoides Miq. Canthium leytense Merr. Canthium libericum Dinkl. Canthium longipes Geddes Canthium lucidum R.Br. Canthium macrocarpum Thwaites Canthium megacarpum Merr. Canthium megistocarpum Merr. & L.M.Perry Canthium merrillianum Mabb. Canthium merrillii Christoph. Canthium mite Bartl. ex DC. Canthium molle King & Gamble Canthium moluccanum Roxb. Canthium neilgherrense Wight Canthium oblongifolium Quisumb. & Merr. Canthium oblongum Kaneh. Canthium oliganthum Boerl. Canthium oligocarpum Hiern Canthium parvifolium Roxb. Canthium paucinervium Merr. Canthium pedunculare Cav. Canthium perakanthus ined. Canthium polyanthum Miq. Canthium puberulum Thwaites ex Hook.f. Canthium quadratum Craib Canthium rheedei DC. Canthium sarcocarpum Merr. Canthium sarmentosum Craib Canthium scabridum Ridl. Canthium scandens Blume Canthium schlechterianum Merr. & L.M.Perry Canthium siamense Pit. Canthium simile Merr. & Chun Canthium sordidum Bullock Canthium spinosum Kuntze Canthium spirostylum Miq. Canthium stellulatum Craib Canthium strigosum Craib Canthium strychnoides Craib Canthium subaureum Craib Canthium subcapitatum Merr. Canthium suberosum Codd Canthium subsessilifolium Merr. Canthium sumatranum Miq. Canthium tavoyanum Merr. Canthium travancoricum Bedd. Canthium trichophorum Quisumb. & Merr. Canthium umbelligerum Miq. Canthium vanwykii Tilney & Kok Canthium villarii Vidal Canthium violaceum Zoll. & Moritzi
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