Branta


The black geese of the genus Branta are waterfowl belonging to the true geese and swans subfamily Anserinae. They occur in the northern coastal regions of the Palearctic and all over North America, migrating to more southernly coasts in winter, and as resident birds in the Hawaiian Islands. Alone in the Southern Hemisphere, a self-sustaining feral population derived from introduced Canada geese is also found in New Zealand.
The scientific name Branta is a Latinised form of Old Norse Brandgás, "burnt goose". The black geese derive their vernacular name for the prominent areas of black coloration found in all species. They can be distinguished from all other true geese by their legs and feet, which are black or very dark grey. Furthermore, they have black bills and large areas of black on the head and neck, with white markings that can be used to tell apart most species. As with most geese, their undertail and uppertail coverts are white. They are also on average smaller than other geese, though some very large taxa are known, which rival the swan goose and the black-necked swan in size.
The Eurasian species of black geese have a more coastal distribution compared to the grey geese which share the same general area of occurrence, not being found far inland even in winter. This does not hold true for the American and Pacific species, in whose ranges grey geese are for the most part absent.

Systematics

Phylogeny

Based on the Taxonomy in Flux from John Boyd's website.

Living species and taxonomy

6–8 living species of black geese are known. In addition, one species has been described from subfossil remains found in the Hawaiian Islands, where it became extinct in prehistoric times. Another undescribed prehistoric species from the Big Island of Hawai‘i was extremely large and flightless; it is tentatively assigned to this genus due to being very peculiar. It is fairly certain that at least another species of this genus awaits discovery on the Big Island, judging from the facts that at least one species of Branta was found on every major Hawaiian island, and that remains of such birds have not been intentionally searched for on the Big Island.
The species are:
ImageScientific nameCommon NameDistribution
Branta bernicla – three subspeciesBrant, or brent gooseHigh-Arctic tundra. Circumpolar; several distinct breeding populations, which winter in particular areas along northern temperate-zone Atlantic and Pacific coastlines.
Branta leucopsisBarnacle gooseArctic islands of the Northeast Atlantic
Branta canadensisCanada gooseTemperate regions of North America
Branta hutchinsii – formerly included in B. canadensisCackling gooseNorth America, northern Canada and Alaska
Branta ruficollisRed-breasted gooseArctic Siberia, mainly on the Taymyr Peninsula, with smaller populations in the Gydan and Yamal peninsulas, northwestern shores of the Black Sea in Bulgaria, Romania and Ukraine
Branta sandvicensisNene, nēnē, or Hawaiian gooseHawaiian Islands

Tentatively assigned to Branta:
The relationships of the enigmatic Geochen rhuax were long unresolved. After reexamination of the subfossil material and comparisms with other subfossil bones assigned to the genus Branta it was redescribed as Branta rhuax in 2013. It was another prehistoric Big Island form and remains known only from some parts of a single bird's skeleton, which were much damaged because the bird apparently died in a volcanic eruption, with the bones being found in an ash-filled depression under a lava flow. A presumed relation to the shelducks proposed by Lester Short in 1970 was generally considered highly unlikely due to that group's biogeography, but more recently, bones of a shelduck-like bird have been found on Kaua‘i. Whether this latter anatid was indeed a shelduck is presently undetermined.
Similarly, two bones found on Oʻahu indicate the erstwhile present of a gigantic waterfowl on this island. Its relationships relative to this genus and the moa-nalos, enormous goose-like dabbling ducks, are completely undeterminable at present.

Early fossil record

Several fossil species of Branta have been described. Since the true geese are hardly distinguishable by anatomical features, the allocation of these to this genus is somewhat uncertain. A number of supposed prehistoric grey geese have been described from North America, partially from the same sites as species assigned to Branta. Whether these are correctly assigned – meaning that the genus Anser was once much more widespread than today and that it coexisted with Branta in freshwater habitat which it today does only most rarely – is not clear. Especially in the case of B. dickeyi and B. howardae, doubts have been expressed about its correct generic assignment.
The former "Branta" minuscula is now placed with the prehistoric American shelducks, Anabernicula. On the other hand, a goose fossil from the Early-Middle Pleistocene of El Salvador is highly similar to Anser and given its age and biogeography it is likely to belong to that genus or Branta.

Footnotes