The islands comprising this ecoregion share a long history of isolation, both from other landmasses and each other. The isolation, combined with harsh climates characterised by low temperatures, strong westerly winds and few hours of sunlight in winter, have resulted in the evolution of many endemic plants and animals, though species richness is relatively low. Wind speeds reach an average of while even in summer the thick cloud cover prevents much sunlight from penetrating. The Bounty Islands are small granite rocks, while the small Antipodes Islands group, the largest group the Auckland Islands and Campbell Island are volcanic in origin. Macquarie Island is the furthest south and the coldest. Where present, soils are mainly boggy peats, up to deep in flat areas. None of the islands are inhabited although there are ongoing research projects including a permanent base of the Australian Antarctic Division on Macquarie Island.
Flora
Vegetation may include low forests of Southern rātā in the more sheltered areas of the Aucklands and parts of Campbell Island, with tussock grassland, shrubland, herbfield, feldmark and cushion plants elsewhere. The islands represent a transition zone between the Antarctic to the south and temperate climates to the north. Individual species include many endemics, such as a Cyathea tree fern, which are not found any further south in the world, along with others that also occur in New Zealand and further north. Macquarie Island, being colder, does not sustain any wooded plants, while the small Bounty Islands lack soils and their flora is largely restricted to algae and lichens on the rocks. The islands are home to a number of rare plants, including a unique genus, the Auckland Island Pleurophyllum, and the only subantarctic orchids, Nematoceras dienemum and Nematoceras sulcatum of Macquarie Island.
Although the islands have been partially occupied at various times the habitats remain largely unspoilt. However, introduced animals prey on native wildlife and graze on the plant cover while both mammals and seabirds are vulnerable to entanglement in deep-sea fishing equipment. Longline fishing for tuna and trawling for squid are particularly damaging. Current predators on the islands include rats and cats which are being systematically removed while cautionary procedures have been implemented to prevent more alien species from establishing. All cats have now been removed from Macquarie Island and there appears to have been an immediate increase in some seabird populations. All of the islands are nature reserves, World Heritage Sites and a Centre of Plant Diversity. Permission to land must be obtained from the Department of Conservation for the New Zealand islands and the Tasmania Parks and Wildlife Service for Macquarie.